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brush noseburn, sticky noseburn

betony-leaf noseburn

Habit Subshrubs or vines, 3–10 dm. Herbs or subshrubs, 2–5 dm.
Stems

trailing or twining, dark green, apex flexuous.

erect to trailing, green to whitish green, apex never flexuous.

Leaves

petiole 6–22 mm;

blade narrowly ovate to lanceolate, 2.5–4 × 1.5–2 cm, base shallowly cordate to truncate, margins serrate to crenate, apex acute to acuminate.

petiole 10–40 mm;

blade triangular-lanceolate to triangular-ovate, 1.5–6 × 1–3.5 cm, base cordate to truncate, margins serrate, apex acute.

Inflorescences

terminal (often appearing leaf-opposed), glands stipitate, prominent throughout, staminate flowers 10–30 per raceme;

staminate bracts 0.5–1.5 mm.

terminal (often appearing leaf-opposed), glands absent, staminate flowers 15–80 per raceme, distally clustered;

staminate bracts 1–2 mm.

Pedicels

staminate 1–2 mm, persistent base 0.3–0.7 mm;

pistillate 3–7 mm in fruit.

staminate 0.7–1 mm, persistent base 0.3–0.6 mm, shorter than subtending bract;

pistillate 3–4 mm in fruit.

Staminate flowers

sepals 3, green, 0.7–1.2 mm;

stamens 3, filaments 0.2–0.4 mm.

sepals 3–4, green, sometimes red-tinged, 1.2–2.3 mm;

stamens 3(–4), filaments 0.4–1 mm.

Pistillate flowers

sepals lanceolate, 0.7–1.5 mm;

styles connate 1/3 length;

stigmas smooth to undulate.

sepals lanceolate, 1.8–5 mm;

styles connate 1/3 length;

stigmas papillate.

Capsules

4–5 mm wide.

7–9 mm wide.

Seeds

dark brown to black, 1.9–2.2 mm.

dark brown with light brown streaks, 3–4 mm.

Tragia glanduligera

Tragia betonicifolia

Phenology Flowering late spring; fruiting late summer–fall. Flowering late spring–summer; fruiting summer–fall.
Habitat Dry, sandy limestone soils, abandoned home sites and mesquite scrub. Dry, sandy soils, disturbed fields, prairies, open woods.
Elevation 10–80 m. (0–300 ft.) 0–400 m. (0–1300 ft.)
Distribution
from FNA
TX; e Mexico; s Mexico; Central America (Guatemala)
[BONAP county map]
from FNA
AL; AR; KS; LA; MO; MS; OK; TN; TX; VA
[WildflowerSearch map]
[BONAP county map]
Discussion

Southern Texas is the northernmost distribution of Tragia glanduligera. In Mexico, it is found in tropical deciduous forests in Campeche, Nuevo León, Tabasco, Veracruz, and Yucatan. This species and T. jonesii are the only species in the flora area with stipitate glands on the inflorescence. Tragia glanduligera differs from T. jonesii by its leaf blade margins with 10–15 smaller teeth per side, shorter staminate pedicels, and truncate to weakly cordate leaf blade bases.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants of Tragia betonicifolia resemble those of T. urticifolia but differ in the greater number of branches from the root crowns, the shorter length of the persistent staminate flower pedicel bases, the longer, narrower pistillate sepals, and the distally clustered arrangement of the staminate flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 187. FNA vol. 12, p. 186.
Parent taxa Euphorbiaceae > Tragia Euphorbiaceae > Tragia
Sibling taxa
T. amblyodonta, T. betonicifolia, T. brevispica, T. cordata, T. jonesii, T. laciniata, T. leptophylla, T. nepetifolia, T. nigricans, T. ramosa, T. saxicola, T. smallii, T. urens, T. urticifolia
T. amblyodonta, T. brevispica, T. cordata, T. glanduligera, T. jonesii, T. laciniata, T. leptophylla, T. nepetifolia, T. nigricans, T. ramosa, T. saxicola, T. smallii, T. urens, T. urticifolia
Synonyms T. urticifolia var. texana
Name authority Pax & K. Hoffmann: in H. G. A. Engler, Pflanzenr. 68[IV,147]: 55. (1919) Nuttall: Trans. Amer. Philos. Soc., n. s. 5: 173. (1835) — (as betonicaefolia)
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