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betony-leaf noseburn

fine-leaf noseburn

Habit Herbs or subshrubs, 2–5 dm. Herbs, 1–4.5 dm.
Stems

erect to trailing, green to whitish green, apex never flexuous.

erect, brownish red to maroon-green, apex never flexuous.

Leaves

petiole 10–40 mm;

blade triangular-lanceolate to triangular-ovate, 1.5–6 × 1–3.5 cm, base cordate to truncate, margins serrate, apex acute.

petiole 0.5–2 mm;

blade acicular to narrowly oblong, 1–6 × 0.2–0.6 cm, base acute to subcuneate, margins usually entire, sometimes serrulate, apex acute to obtuse.

Inflorescences

terminal (often appearing leaf-opposed), glands absent, staminate flowers 15–80 per raceme, distally clustered; staminate bracts 1–2 mm.

terminal (appearing leaf opposed) or axillary, glands absent, staminate flowers 2–3(–5) per raceme; staminate bracts 2–2.5 mm.

Pedicels

staminate 0.7–1 mm, persistent base 0.3–0.6 mm, shorter than subtending bract; pistillate 3–4 mm in fruit.

staminate 1–2 mm, persistent base 0.5–0.8 mm, pistillate 2–3 mm in fruit.

Staminate flowers

sepals 3–4, green, sometimes red-tinged, 1.2–2.3 mm;

stamens 3(–4), filaments 0.4–1 mm.

sepals 3–4(–5), green, 1–2.5 mm;

stamens 3–4, filaments 1.2–1.4 mm, connate basally.

Pistillate flowers

sepals lanceolate, 1.8–5 mm;

styles connate 1/3 length;

stigmas papillate.

sepals lanceolate, 1.5–2 mm;

styles connate 1/5 length;

stigmas undulate to slightly papillate.

Capsules

7–9 mm wide.

4–5 mm wide.

Seeds

dark brown with light brown streaks, 3–4 mm.

mottled dark olive brown, 2.5–3 mm.

Tragia betonicifolia

Tragia leptophylla

Phenology Flowering late spring–summer; fruiting summer–fall. Flowering spring–fall; fruiting late spring and fall.
Habitat Dry, sandy soils, disturbed fields, prairies, open woods. Dry streams and river margins with limestone cobble substrates.
Elevation 0–400 m. (0–1300 ft.) 400–700 m. (1300–2300 ft.)
Distribution
from FNA
AL; AR; KS; LA; MO; MS; OK; TN; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

Plants of Tragia betonicifolia resemble those of T. urticifolia but differ in the greater number of branches from the root crowns, the shorter length of the persistent staminate flower pedicel bases, the longer, narrower pistillate sepals, and the distally clustered arrangement of the staminate flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Tragia leptophylla is known from the western part of the Edwards Plateau region in west-central Texas. K. I. Miller and G. L. Webster (1967) did not recognize T. leptophylla as a distinct species; they treated it as a synonym of T. ramosa. Tragia leptophylla differs from T. ramosa in its less branching habit, dark reddish stems, usually entire leaf blade margins, fewer staminate flowers per inflorescence, and riparian limestone cobble habitat.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 186. FNA vol. 12, p. 188.
Parent taxa Euphorbiaceae > Tragia Euphorbiaceae > Tragia
Sibling taxa
T. amblyodonta, T. brevispica, T. cordata, T. glanduligera, T. jonesii, T. laciniata, T. leptophylla, T. nepetifolia, T. nigricans, T. ramosa, T. saxicola, T. smallii, T. urens, T. urticifolia
T. amblyodonta, T. betonicifolia, T. brevispica, T. cordata, T. glanduligera, T. jonesii, T. laciniata, T. nepetifolia, T. nigricans, T. ramosa, T. saxicola, T. smallii, T. urens, T. urticifolia
Synonyms T. urticifolia var. texana T. ramosa var. leptophylla, T. stylaris var. leptophylla
Name authority Nuttall: Trans. Amer. Philos. Soc., n. s. 5: 173. (1835) — (as betonicaefolia) (Torrey) I. M. Johnston: Contr. Gray Herb. 68: 91. (1923)
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