Tragia amblyodonta |
Tragia |
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blunt-tooth noseburn, dog-tooth noseburn |
noseburn |
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Habit | Subshrubs, 1.2–5 dm. | Herbs, subshrubs, or vines, perennial, monoecious [dioecious]; hairy, hairs unbranched, always some stinging (sometimes inconspicuous except on ovaries and capsules), sometimes glandular; latex absent. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | erect to trailing, gray-green, apex often flexuous. |
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Leaves | petiole 4–20(–30) mm; blade usually triangular to subhastate, sometimes ovate, 1–4.5 × 0.8–3 cm, base cordate, hastate, or truncate, margins crenate to serrate, apex acute to obtuse. |
deciduous, alternate, simple (usually 3-foliolate in T. laciniata); stipules present, persistent; petiole present, glands absent; blade usually unlobed, sometimes lobed basally (sometimes deeply 3-lobed in T. laciniata) [palmately lobed], margins serrate, crenate, dentate, or entire, laminar glands absent; venation pinnate or palmate at base, pinnate distally [palmate]. |
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Inflorescences | terminal or axillary, glands absent, staminate flowers 5–16 per raceme; staminate bracts 0.9–2 mm. |
bisexual (pistillate flowers proximal, staminate distal) [unisexual], axillary, terminal, or leaf-opposed, racemes [rarely with single pistillate branch]; glands subtending each bract 0. |
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Pedicels | staminate 0.7–1.2 mm, persistent base 0.2–0.8 mm; pistillate 1.5–4 mm in fruit. |
present, staminate with persistent base, pistillate elongated in fruit. |
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Staminate flowers | sepals 3–4, green, 0.9–1.2 mm; stamens 3–4, filaments 0.2–0.7 mm. |
sepals 3–5, usually green, sometimes reddish green, not petaloid, valvate, distinct; petals 0; nectary absent [present]; stamens 2–6(–10)[–25], distinct or connate basally (connate 1/2 length in T. nigricans); pistillode present [absent]. |
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Pistillate flowers | sepals lanceolate, 1–2.5 mm; styles connate to 1/3 length, short-exserted; stigmas undulate to subpapillate. |
sepals 6, usually green, sometimes reddish green, not petaloid, connate basally; petals 0; nectary absent; pistil 3-carpellate; styles 3, connate basally to 1/2 [most of] length, unbranched. |
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Fruits | capsules, usually 3 carpels maturing, except often 1 maturing in T. brevispica. |
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Capsules | 7–8 mm wide. |
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Seeds | brown with tan mottling, 2.5–3.5 mm. |
globose to ovoid; caruncle absent. |
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2n | = 110. |
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Tragia amblyodonta |
Tragia |
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Phenology | Flowering spring–fall; fruiting summer–late fall. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Dry, rocky, exposed slopes in xerophytic scrub. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 10–1400 m. (0–4600 ft.) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
AZ; NM; TX; Mexico (Chihuahua, Coahuila, Durango, Nuevo León, Tamaulipas) |
United States; Mexico; Central America; South America; West Indies; Asia; Africa; Australia; primarily tropical and subtropical regions |
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Discussion | Tragia amblyodonta is easily distinguished from other members of Tragia by the combination of usually triangular to subhastate leaf blades, gray-green coloration, and painfully stinging hairs. Both stomata diameter and pollen grain size of T. amblyodonta are larger than in any other North American species of Tragia (K. I. Miller and G. L. Webster 1967). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 175 (15 in the flora). Tragia is a taxonomically difficult genus that is characterized by stinging hairs. Although many species of Tragia are twining vines, most species in the flora area are subshrubs or herbs. Some species are used medicinally for their anti-inflammatory, analgesic, vermifugic, and antihyperglycemic properties. Two sections are represented in the flora area: Tragia and Leptobotrys (Baillon) Müller Arg. Molecular phylogenetic analysis (W. M. Cardinal-McTeague and L. J. Gillespie, unpubl.) suggests that Tragia is polyphyletic and that sect. Leptobotrys (T. smallii, T. urens) should be segregated as a distinct genus; these results are supported by pollen morphology (L. J. Gillespie 1994). Tragia volubilis Linnaeus was collected from Florida once (1842–1848, F. Rugel, US), but has not been collected there since and is presumed extirpated in the flora area. This species is widespread in the Caribbean and Latin America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 186. | FNA vol. 12, p. 184. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | T. nepetifolia var. amblyodonta | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Müller Arg.) Pax & K. Hoffmann: in H. G. A. Engler, Pflanzenr. 68[IV,147]: 51. (1919) | Linnaeus: Sp. Pl. 2: 980. (1753): Gen. Pl. ed. 5, 421. (1754) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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