Tragia |
Tragia cordata |
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noseburn |
heart-leaf noseburn |
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Habit | Herbs, subshrubs, or vines, perennial, monoecious [dioecious]; hairy, hairs unbranched, always some stinging (sometimes inconspicuous except on ovaries and capsules), sometimes glandular; latex absent. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | usually decumbent or twining, rarely erect, gray-green to light green, apex flexuous. |
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Leaves | deciduous, alternate, simple (usually 3-foliolate in T. laciniata); stipules present, persistent; petiole present, glands absent; blade usually unlobed, sometimes lobed basally (sometimes deeply 3-lobed in T. laciniata) [palmately lobed], margins serrate, crenate, dentate, or entire, laminar glands absent; venation pinnate or palmate at base, pinnate distally [palmate]. |
petiole 15–85 mm; blade ovate to broadly cordate, 4.5–10(–13) × 3.5–10 cm, base cordate, margins serrate, apex acuminate. |
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Inflorescences | bisexual (pistillate flowers proximal, staminate distal) [unisexual], axillary, terminal, or leaf-opposed, racemes [rarely with single pistillate branch]; glands subtending each bract 0. |
terminal (often appearing leaf-opposed), glands absent, staminate flowers 20–60 per raceme; staminate bracts 1.5–2 mm. |
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Pedicels | present, staminate with persistent base, pistillate elongated in fruit. |
staminate 1.5–2.2 mm, persistent base 0.7–1 mm; pistillate 2.5–3 mm in fruit. |
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Staminate flowers | sepals 3–5, usually green, sometimes reddish green, not petaloid, valvate, distinct; petals 0; nectary absent [present]; stamens 2–6(–10)[–25], distinct or connate basally (connate 1/2 length in T. nigricans); pistillode present [absent]. |
sepals 3, green, 0.7–1 mm; stamens 3, filaments 0.2–0.5 mm. |
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Pistillate flowers | sepals 6, usually green, sometimes reddish green, not petaloid, connate basally; petals 0; nectary absent; pistil 3-carpellate; styles 3, connate basally to 1/2 [most of] length, unbranched. |
sepals elliptic to ovate, 1.5–2 mm; styles connate 1/4–1/3 length; stigmas papillate. |
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Fruits | capsules, usually 3 carpels maturing, except often 1 maturing in T. brevispica. |
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Capsules | 11–13 mm wide. |
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Seeds | globose to ovoid; caruncle absent. |
dark brown, 4.3–5.3 mm. |
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Vines | , 15–20 dm. |
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Tragia |
Tragia cordata |
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Phenology | Flowering spring–fall; fruiting summer–late fall. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Rich deciduous forests, riverbanks, rocky thickets. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 50–500 m. (200–1600 ft.) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
United States; Mexico; Central America; South America; West Indies; Asia; Africa; Australia; primarily tropical and subtropical regions |
AL; AR; FL; GA; IL; IN; KY; LA; MO; MS; OK; TN; TX
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Discussion | Species ca. 175 (15 in the flora). Tragia is a taxonomically difficult genus that is characterized by stinging hairs. Although many species of Tragia are twining vines, most species in the flora area are subshrubs or herbs. Some species are used medicinally for their anti-inflammatory, analgesic, vermifugic, and antihyperglycemic properties. Two sections are represented in the flora area: Tragia and Leptobotrys (Baillon) Müller Arg. Molecular phylogenetic analysis (W. M. Cardinal-McTeague and L. J. Gillespie, unpubl.) suggests that Tragia is polyphyletic and that sect. Leptobotrys (T. smallii, T. urens) should be segregated as a distinct genus; these results are supported by pollen morphology (L. J. Gillespie 1994). Tragia volubilis Linnaeus was collected from Florida once (1842–1848, F. Rugel, US), but has not been collected there since and is presumed extirpated in the flora area. This species is widespread in the Caribbean and Latin America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Both the morphology and ecology of Tragia cordata make it unique among American members of Tragia. The relatively large, heart-shaped leaves separate it from the other Tragia in the flora area; it is the only twining species of Tragia found in the deciduous forest of the Midwest. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 184. | FNA vol. 12, p. 187. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Name authority | Linnaeus: Sp. Pl. 2: 980. (1753): Gen. Pl. ed. 5, 421. (1754) | Michaux: Fl. Bor.-Amer. 2: 176. (1803) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |