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tortula moss, wall screw-moss

desmatodon moss

Leaves

ovate or elliptic to obovate, apex broadly acute to rounded or emarginate, short- to long-awned, margins narrowly recurved or revolute to near apex, commonly bordered distally with 2–4 rows of thicker-walled, papillose to smooth cells;

costa long-excurrent, lacking an adaxial pad of cells, distally narrow, 3–4(–6) cells across the convex adaxial surface;

distal laminal cells hexagonal, 10–15 µm wide, 1:1, strongly papillose with 2(–4)-fid papillae.

obovate to occasionally ovate, apex acute or broadly acute, awned, margins recurved or plane proximally, not bordered;

costa excurrent into the awn, lacking an adaxial pad of cells, distally narrow, 2–3 cells across the convex adaxial surface;

distal laminal cells hexagonal to rhomboidal, width 15–22 µm wide, 1:1 near apex but elongate at mid leaf and basally, smooth.

Seta

0.6–1.5 cm.

0.8–1.1 cm.

Sexual condition

autoicous.

autoicous.

Capsule

stegocarpic, not systylius, cylindric, erect and nearly straight, urn 1.5–2.7 mm;

peristome 300 µm, teeth of 32 filaments twisted 1/2–2 times, basal membrane low, to 50 µm;

operculum 0.6–1 mm.

systylius, short-cylindric, erect and nearly straight, urn 1.5–2 mm;

peristome 200–250 µm, teeth 16, straight, divided to near base but variously fused or perforate, basal membrane low;

operculum 0.5–0.7 mm.

Spores

8–12 µm, spheric, finely papillose or essentially smooth.

22–30(–35) µm, spheric, warty papillose.

Sporophytes

exerted.

exerted.

Tortula muralis

Tortula systylia

Phenology Capsules mature spring–summer. Capsules mature summer.
Habitat Calcareous rock, often on bricks or walls Soil in rock crevices, tundra
Elevation low to moderate elevations high elevations (0–3700 m) (high elevations (0–12100 ft))
Distribution
from FNA
AK; AL; AR; AZ; CA; CO; DC; FL; GA; IA; KY; LA; MD; MI; MO; NC; ND; NJ; NV; NY; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; BC; NF; NU; ON; West Indies; s South America; Europe; Asia; Africa; Atlantic Islands; Pacific Islands (New Zealand); Australia
[WildflowerSearch map]
from FNA
AK; CA; CO; WY; AB; BC; MB; NF; NT; NU; QC; YT; Greenland; Europe; Asia
Discussion

Tortula muralis, T. brevipes, and T. plinthobia form an apparently intergrading cline in sporophyte characters, sexual condition, and elaboration of the leaf border. There may be evolutionary advantages associated with such variability, but specimens are occasionally difficult to name satisfactorily. The laminal border of 2–4 rows of thicker walled cells is usually hidden in the margin recurvature except at the leaf apex.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Tortula systylia may be confused with Stegonia but the latter has a distinct triangle of echlorophyllose cells at the apex of the much broader leaves. It is similar also to T. hoppeana, but the laminal cells are smooth. After dehiscence, the systylius capsules retain a raised operculum on the columellar stalk.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 595. FNA vol. 27, p. 599.
Parent taxa Pottiaceae > subfam. Pottioideae > Tortula Pottiaceae > subfam. Pottioideae > Tortula
Sibling taxa
T. acaulon, T. amplexa, T. atrovirens, T. bolanderi, T. brevipes, T. californica, T. cernua, T. cuneifolia, T. deciduidentata, T. guepinii, T. hoppeana, T. inermis, T. lanceola, T. laureri, T. leucostoma, T. modica, T. mucronifolia, T. nevadensis, T. obtusifolia, T. plinthobia, T. porteri, T. protobryoides, T. subulata, T. systylia, T. truncata
T. acaulon, T. amplexa, T. atrovirens, T. bolanderi, T. brevipes, T. californica, T. cernua, T. cuneifolia, T. deciduidentata, T. guepinii, T. hoppeana, T. inermis, T. lanceola, T. laureri, T. leucostoma, T. modica, T. mucronifolia, T. muralis, T. nevadensis, T. obtusifolia, T. plinthobia, T. porteri, T. protobryoides, T. subulata, T. truncata
Synonyms Desmatodon systylius
Name authority Hedwig: Sp. Musc. Frond., 123. (1801) (Schimper) Lindberg: Musc. Scand., 20. (1879)
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