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tortula moss, wall screw-moss

obtuseleaf desmatodon moss

Leaves

ovate or elliptic to obovate, apex broadly acute to rounded or emarginate, short- to long-awned, margins narrowly recurved or revolute to near apex, commonly bordered distally with 2–4 rows of thicker-walled, papillose to smooth cells;

costa long-excurrent, lacking an adaxial pad of cells, distally narrow, 3–4(–6) cells across the convex adaxial surface;

distal laminal cells hexagonal, 10–15 µm wide, 1:1, strongly papillose with 2(–4)-fid papillae.

ovate to ligulate, apex rounded or occasionally broadly acute, apiculate, margins strongly recurved or revolute in distal 4/5 of leaf, sometimes bordered distally with (2–)4 rows of thicker-walled cells, or occasionally border of rhomboidal, smooth cells commonly hidden in recurvature;

costa ending in apiculus or occasionally percurrent or subpercurrent, lacking an adaxial pad of swollen cells or this poorly developed, distally not broadening, to 5–6 cells across the convex adaxial surface;

distal laminal cells hexagonal or subquadrate, 10–13 µm wide, 1:1, strongly papillose with 2-fid or multiplex papillae.

Seta

0.6–1.5 cm.

0.5–0.8 cm.

Sexual condition

autoicous.

autoicous, but plants often bearing only antheridia or archegonia.

Capsule

stegocarpic, not systylius, cylindric, erect and nearly straight, urn 1.5–2.7 mm;

peristome 300 µm, teeth of 32 filaments twisted 1/2–2 times, basal membrane low, to 50 µm;

operculum 0.6–1 mm.

stegocarpic, cylindric or ovate, erect and nearly straight, urn 1–2.5 mm;

peristome rudimentary or to 500 µm, teeth 16, irregular, divided to base and filamentous when long, straight or twisted, basal membrane very low or absent;

operculum 0.7–1 mm.

Spores

8–12 µm, spheric, finely papillose or essentially smooth.

9–11(–15) µm, spheric, finely papillose or essentially smooth.

Sporophytes

exerted.

exerted.

Tortula muralis

Tortula obtusifolia

Phenology Capsules mature spring–summer. Capsules mature spring–fall.
Habitat Calcareous rock, often on bricks or walls Soil, rock, limestone, calcareous sandstone, stone walls, crevices, ledges
Elevation low to moderate elevations low to high elevations (0-3300 m) (low to high elevations (0-10800 ft))
Distribution
from FNA
AK; AL; AR; AZ; CA; CO; DC; FL; GA; IA; KY; LA; MD; MI; MO; NC; ND; NJ; NV; NY; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; BC; NF; NU; ON; West Indies; s South America; Europe; Asia; Africa; Atlantic Islands; Pacific Islands (New Zealand); Australia
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from FNA
AK; AZ; CA; CO; GA; IA; IN; KS; MI; MN; MO; MT; NE; NJ; NM; OH; OK; PA; SD; TN; TX; UT; WA; WI; WY; AB; BC; MB; NB; ON; QC; Europe; Asia; Africa
[WildflowerSearch map]
Discussion

Tortula muralis, T. brevipes, and T. plinthobia form an apparently intergrading cline in sporophyte characters, sexual condition, and elaboration of the leaf border. There may be evolutionary advantages associated with such variability, but specimens are occasionally difficult to name satisfactorily. The laminal border of 2–4 rows of thicker walled cells is usually hidden in the margin recurvature except at the leaf apex.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The variation in peristome development of Tortula obtusifolia is considerable, and matches intraspecifically the variation between the peristomes of species previously recognized in the segregate genus Desmatodon and in Tortula in the strict sense. This species may be confused with T. atrovirens but the latter has a massive pad of bulging cells on the costa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 595. FNA vol. 27, p. 597.
Parent taxa Pottiaceae > subfam. Pottioideae > Tortula Pottiaceae > subfam. Pottioideae > Tortula
Sibling taxa
T. acaulon, T. amplexa, T. atrovirens, T. bolanderi, T. brevipes, T. californica, T. cernua, T. cuneifolia, T. deciduidentata, T. guepinii, T. hoppeana, T. inermis, T. lanceola, T. laureri, T. leucostoma, T. modica, T. mucronifolia, T. nevadensis, T. obtusifolia, T. plinthobia, T. porteri, T. protobryoides, T. subulata, T. systylia, T. truncata
T. acaulon, T. amplexa, T. atrovirens, T. bolanderi, T. brevipes, T. californica, T. cernua, T. cuneifolia, T. deciduidentata, T. guepinii, T. hoppeana, T. inermis, T. lanceola, T. laureri, T. leucostoma, T. modica, T. mucronifolia, T. muralis, T. nevadensis, T. plinthobia, T. porteri, T. protobryoides, T. subulata, T. systylia, T. truncata
Synonyms Barbula obtusifolia, Desmatodon arenaceus, Desmatodon coloradensis, Desmatodon obtusifolius
Name authority Hedwig: Sp. Musc. Frond., 123. (1801) (Schwägrichen) Mathieu: Fl. Gén. Belgique 2: 48. (1853)
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