Tortula muralis |
Tortula lanceola |
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tortula moss, wall screw-moss |
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Leaves | ovate or elliptic to obovate, apex broadly acute to rounded or emarginate, short- to long-awned, margins narrowly recurved or revolute to near apex, commonly bordered distally with 2–4 rows of thicker-walled, papillose to smooth cells; costa long-excurrent, lacking an adaxial pad of cells, distally narrow, 3–4(–6) cells across the convex adaxial surface; distal laminal cells hexagonal, 10–15 µm wide, 1:1, strongly papillose with 2(–4)-fid papillae. |
obovate, elliptic or less commonly ovate, apex broadly acute to occasionally rounded, awned or long-mucronate, margins strongly and broadly recurved in proximal 4/5 of leaf, not bordered; costa long-excurrent, with an elongate adaxial pad of bulging, rounded cells with 3–4 simple papillae, distally narrow, 2–4 cells across adaxial surface; distal laminal cells subquadrate, 12–16 µm wide, 1:1, smooth or less commonly papillose with 1(–2) scattered simple papillae. |
Seta | 0.6–1.5 cm. |
ca. 0.4(–0.9) cm. |
Sexual condition | autoicous. |
autoicous. |
Capsule | stegocarpic, not systylius, cylindric, erect and nearly straight, urn 1.5–2.7 mm; peristome 300 µm, teeth of 32 filaments twisted 1/2–2 times, basal membrane low, to 50 µm; operculum 0.6–1 mm. |
stegocarpic, not systylius, elliptic or cylindric, or erect and nearly straight, urn ca. 0.8–1.2 mm; peristome 100–150 µm, teeth 16, ligulate straight, or short, rudimentary, basal membrane lacking or very low; operculum 0.5 mm. |
Spores | 8–12 µm, spheric, finely papillose or essentially smooth. |
15–18 µm, spheric, finely papillose. |
Sporophytes | exerted. |
exerted. |
Tortula muralis |
Tortula lanceola |
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Phenology | Capsules mature spring–summer. | Capsules mature spring. |
Habitat | Calcareous rock, often on bricks or walls | Soil or walls, often in calcareous regions |
Elevation | low to moderate elevations | low to moderate elevations |
Distribution |
AK; AL; AR; AZ; CA; CO; DC; FL; GA; IA; KY; LA; MD; MI; MO; NC; ND; NJ; NV; NY; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; BC; NF; NU; ON; West Indies; s South America; Europe; Asia; Africa; Atlantic Islands; Pacific Islands (New Zealand); Australia
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AZ; ID; SC; TX; Europe; Asia; n Africa |
Discussion | Tortula muralis, T. brevipes, and T. plinthobia form an apparently intergrading cline in sporophyte characters, sexual condition, and elaboration of the leaf border. There may be evolutionary advantages associated with such variability, but specimens are occasionally difficult to name satisfactorily. The laminal border of 2–4 rows of thicker walled cells is usually hidden in the margin recurvature except at the leaf apex. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The tiny but often fruiting Tortula lanceola is uncommon in the flora area. The bulging, rounded cells of the adaxial costal pad have the distinctive appearance of cows’ udders, and viewed laterally have the appearance of longitudinal ridges. H. A. Crum and L. E. Anderson (1981) pointed out that the antheridia are borne naked in axillary clusters, but occasionally bracts, albeit rudimentary, are present. Tortula acaulon has much the same gametophyte, but the sporophyte is reduced and cleistocarpic. Tortula atrovirens and Crossidium aberrans have a somewhat similar leaf section but the former lacks an awn, and the latter has a short awn, and the adaxial costal pad is of low 1–2-celled filaments. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 595. | FNA vol. 27, p. 592. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Tortula | Pottiaceae > subfam. Pottioideae > Tortula |
Sibling taxa | ||
Synonyms | Encalypta lanceolata, Pottia lanceolata | |
Name authority | Hedwig: Sp. Musc. Frond., 123. (1801) | R. H. Zander: Bull. Buffalo Soc. Nat. Sci. 32: 223. (1993) |
Web links |