Tortula muralis |
Tortula inermis |
|
---|---|---|
tortula moss, wall screw-moss |
tortula moss |
|
Leaves | ovate or elliptic to obovate, apex broadly acute to rounded or emarginate, short- to long-awned, margins narrowly recurved or revolute to near apex, commonly bordered distally with 2–4 rows of thicker-walled, papillose to smooth cells; costa long-excurrent, lacking an adaxial pad of cells, distally narrow, 3–4(–6) cells across the convex adaxial surface; distal laminal cells hexagonal, 10–15 µm wide, 1:1, strongly papillose with 2(–4)-fid papillae. |
lingulate, apex rounded-acute, apiculate to short-mucronate, margins recurved from base to near the apex, not bordered; costa subpercurrent, percurrent or very short-excurrent, lacking an adaxial pad of cells but distally narrowing or weakly thickened, 3–4(–5) cells across the convex adaxial surface; distal laminal cells hexagonal, (12–)15–18 µm wide, 1:1, strongly papillose with 4–5 2-fid papillae. |
Seta | 0.6–1.5 cm. |
1.2–1.5(–2.5) cm. |
Sexual condition | autoicous. |
gonioautoicous. |
Capsule | stegocarpic, not systylius, cylindric, erect and nearly straight, urn 1.5–2.7 mm; peristome 300 µm, teeth of 32 filaments twisted 1/2–2 times, basal membrane low, to 50 µm; operculum 0.6–1 mm. |
stegocarpic, not systylius, cylindric, erect and nearly straight, urn usually 3–4 mm; peristome length 1000–1500 µm, teeth of 32 filaments twisted at least one full turn, basal membrane 300–600 µm; operculum 1.3–1.6(–2) mm. |
Spores | 8–12 µm, spheric, finely papillose or essentially smooth. |
11–15 µm, spheric, finely papillose. |
Sporophytes | exerted. |
exerted. |
Tortula muralis |
Tortula inermis |
|
Phenology | Capsules mature spring–summer. | Capsules mature spring. |
Habitat | Calcareous rock, often on bricks or walls | Soil, rock |
Elevation | low to moderate elevations | low to high elevations (0-2000 m) (low to high elevations (0-6600 ft)) |
Distribution |
AK; AL; AR; AZ; CA; CO; DC; FL; GA; IA; KY; LA; MD; MI; MO; NC; ND; NJ; NV; NY; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; BC; NF; NU; ON; West Indies; s South America; Europe; Asia; Africa; Atlantic Islands; Pacific Islands (New Zealand); Australia
|
AZ; CA; CO; NM; NV; SD; TX; UT; WY; Mexico (Baja California, Chihuahua, Sonora); Europe; s Asia; n Africa |
Discussion | Tortula muralis, T. brevipes, and T. plinthobia form an apparently intergrading cline in sporophyte characters, sexual condition, and elaboration of the leaf border. There may be evolutionary advantages associated with such variability, but specimens are occasionally difficult to name satisfactorily. The laminal border of 2–4 rows of thicker walled cells is usually hidden in the margin recurvature except at the leaf apex. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Tortula inermis has the aspect of a Syntrichia with its ligulate, apiculate leaves and strong costa, but the plant is yellow or orange in KOH solution, and the costal section reveals a rounded stereid band. It is related to T. subulata and T. mucronifolia but the lack of a strong mucro and the narrow but nearly complete recurving of the leaf margins are diagnostic. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 595. | FNA vol. 27, p. 600. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Tortula | Pottiaceae > subfam. Pottioideae > Tortula |
Sibling taxa | ||
Synonyms | Syntrichia subulata var. inermis, Syntrichia inermis | |
Name authority | Hedwig: Sp. Musc. Frond., 123. (1801) | (Bridel) Montagne: Arch. Bot. (Leipzig) 1: 136. (1832) |
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