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tortula moss, wall screw-moss

Leaves

ovate or elliptic to obovate, apex broadly acute to rounded or emarginate, short- to long-awned, margins narrowly recurved or revolute to near apex, commonly bordered distally with 2–4 rows of thicker-walled, papillose to smooth cells;

costa long-excurrent, lacking an adaxial pad of cells, distally narrow, 3–4(–6) cells across the convex adaxial surface;

distal laminal cells hexagonal, 10–15 µm wide, 1:1, strongly papillose with 2(–4)-fid papillae.

elliptic, apex broadly acute to rounded-acute, mucronate or apiculate, margins recurved at mid leaf, bordered throughout or in lower 2/3 with 2(–3) rows of narrower, short- to long-rectangular, thicker-walled cells that are 1–2-stratose;

costa percurrent or short-excurrent, lacking an adaxial pad of cells, distally narrow, 3–4 cells across the convex adaxial surface;

distal laminal cells hexagonal to rectangular, (13–)16–24(–28) µm wide, 1–2:1, not or rarely very weakly papillose.

Seta

0.6–1.5 cm.

0.6–1.2 cm.

Sexual condition

autoicous.

autoicous.

Capsule

stegocarpic, not systylius, cylindric, erect and nearly straight, urn 1.5–2.7 mm;

peristome 300 µm, teeth of 32 filaments twisted 1/2–2 times, basal membrane low, to 50 µm;

operculum 0.6–1 mm.

stegocarpic, not systylius, short-cylindric and curved or ovoid and bilaterally symmetrical, nodding or occasionally nearly horizontal, urn 1–1.8 mm;

peristome weak, ca. 100–200 µm, teeth 16, not twisted, divided nearly to the base into 2–3 branches and variously perforate, basal membrane very low or absent, or rudimentary or fragmentary and attached to the inside of the operculum;

operculum 0.2–1.5 mm.

Spores

8–12 µm, spheric, finely papillose or essentially smooth.

25–35 µm, spheric, densely papillose.

Sporophytes

exerted.

exerted.

Tortula muralis

Tortula cernua

Phenology Capsules mature spring–summer. Capsules mature summer.
Habitat Calcareous rock, often on bricks or walls Soil, limestone
Elevation low to moderate elevations low to moderate elevations
Distribution
from FNA
AK; AL; AR; AZ; CA; CO; DC; FL; GA; IA; KY; LA; MD; MI; MO; NC; ND; NJ; NV; NY; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; BC; NF; NU; ON; West Indies; s South America; Europe; Asia; Africa; Atlantic Islands; Pacific Islands (New Zealand); Australia
[WildflowerSearch map]
from FNA
AK; AZ; CO; IA; IL; ME; MT; ND; NV; OH; UT; WI; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; QC; SK; YT; Greenland; Europe; Asia
Discussion

Tortula muralis, T. brevipes, and T. plinthobia form an apparently intergrading cline in sporophyte characters, sexual condition, and elaboration of the leaf border. There may be evolutionary advantages associated with such variability, but specimens are occasionally difficult to name satisfactorily. The laminal border of 2–4 rows of thicker walled cells is usually hidden in the margin recurvature except at the leaf apex.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Tortula cernua differs from T. laureri in the usually smooth laminal cells, the mostly ovoid and bilaterally symmetrical capsule inclined to horizontal, and peristome not twisted. Its range is more southerly and at lower elevations. The small and fragile peristome commonly falls with the operculum, being attached to its inner surface, and the capsule may appear gymnostomous. Given that the uncommon T. randii is only distinguished from T. cernua by the apparent absence of a peristome and has much the same range, and all specimens of that species I have seen are indeed peristomate or, as in A. J. Grout, N. Amer. Musci Perf. 197, MO, have smooth fragments of a peristome attached to the inner surface of the operculum, I agree with J. M. Holzinger (1925) that it should not be recognized as distinct.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 595. FNA vol. 27, p. 601.
Parent taxa Pottiaceae > subfam. Pottioideae > Tortula Pottiaceae > subfam. Pottioideae > Tortula
Sibling taxa
T. acaulon, T. amplexa, T. atrovirens, T. bolanderi, T. brevipes, T. californica, T. cernua, T. cuneifolia, T. deciduidentata, T. guepinii, T. hoppeana, T. inermis, T. lanceola, T. laureri, T. leucostoma, T. modica, T. mucronifolia, T. nevadensis, T. obtusifolia, T. plinthobia, T. porteri, T. protobryoides, T. subulata, T. systylia, T. truncata
T. acaulon, T. amplexa, T. atrovirens, T. bolanderi, T. brevipes, T. californica, T. cuneifolia, T. deciduidentata, T. guepinii, T. hoppeana, T. inermis, T. lanceola, T. laureri, T. leucostoma, T. modica, T. mucronifolia, T. muralis, T. nevadensis, T. obtusifolia, T. plinthobia, T. porteri, T. protobryoides, T. subulata, T. systylia, T. truncata
Synonyms Dermatodon cernuus, Dermatodon randii, Pottia randii, T. randii
Name authority Hedwig: Sp. Musc. Frond., 123. (1801) (Huebener) Lindberg: Musc. Scand., 20. (1879)
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