Tortula muralis |
Tortula atrovirens |
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tortula moss, wall screw-moss |
convolute desmatodon moss |
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Leaves | ovate or elliptic to obovate, apex broadly acute to rounded or emarginate, short- to long-awned, margins narrowly recurved or revolute to near apex, commonly bordered distally with 2–4 rows of thicker-walled, papillose to smooth cells; costa long-excurrent, lacking an adaxial pad of cells, distally narrow, 3–4(–6) cells across the convex adaxial surface; distal laminal cells hexagonal, 10–15 µm wide, 1:1, strongly papillose with 2(–4)-fid papillae. |
ovate to lingulate, apex broadly acute to rounded, apiculate or mucronate, margins revolute or occasionally plane, often bordered proximally with rectangular cells usually hidden in the revolute margins; costa percurrent or ending in apiculus or short mucro, with an adaxial pad of swollen cells similar to those of the lamina, distally broad, (4–)6(–8) cells across adaxial surface; distal laminal cells subquadrate, 9–13(–18) µm wide, µm, 1:1; strongly papillose. |
Seta | 0.6–1.5 cm. |
0.6–1.2 cm. |
Sexual condition | autoicous. |
autoicous. |
Capsule | stegocarpic, not systylius, cylindric, erect and nearly straight, urn 1.5–2.7 mm; peristome 300 µm, teeth of 32 filaments twisted 1/2–2 times, basal membrane low, to 50 µm; operculum 0.6–1 mm. |
stegocarpic, not systylius, cylindric to ellipsoidal, erect and nearly straight, urn 0.9–1.5 mm; peristome commonly 210–390 µm, teeth of 32 filaments, weakly twisted, occasionally rudimentary, basal membrane 40–50 µm; operculum 0.5–0.7 mm. |
Spores | 8–12 µm, spheric, finely papillose or essentially smooth. |
15–18 µm, broadly ellipsoidal, nearly smooth. |
Sporophytes | exerted. |
exerted. |
Tortula muralis |
Tortula atrovirens |
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Phenology | Capsules mature spring–summer. | Capsules mature winter–spring. |
Habitat | Calcareous rock, often on bricks or walls | Exposed soil, volcanic ash, rock, often calcareous |
Elevation | low to moderate elevations | low to high elevations (0-2200 m) (low to high elevations (0-7200 ft)) |
Distribution |
AK; AL; AR; AZ; CA; CO; DC; FL; GA; IA; KY; LA; MD; MI; MO; NC; ND; NJ; NV; NY; OK; OR; PA; SC; TN; TX; UT; VA; WA; WV; BC; NF; NU; ON; West Indies; s South America; Europe; Asia; Africa; Atlantic Islands; Pacific Islands (New Zealand); Australia
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AZ; CA; CO; ID; MN; NM; NV; TX; UT; AB; BC; Mexico; South America; Europe; Asia; Africa; Atlantic Islands; Pacific Islands (Hawaii, New Zealand); Australia
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Discussion | Tortula muralis, T. brevipes, and T. plinthobia form an apparently intergrading cline in sporophyte characters, sexual condition, and elaboration of the leaf border. There may be evolutionary advantages associated with such variability, but specimens are occasionally difficult to name satisfactorily. The laminal border of 2–4 rows of thicker walled cells is usually hidden in the margin recurvature except at the leaf apex. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Tortula atrovirens has a massive pad of bulging cells covering the costa, and may be mistaken for a Crossidium, which, however, has a ventral pad of filaments. It is somewhat similar to T. obtusifolia, which also has weakly differentiated, smooth, somewhat elongate marginal cells, but that species of more mesic climates and more northern latitudes has spatulate leaves lacking an adaxial pad. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 595. | FNA vol. 27, p. 602. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Tortula | Pottiaceae > subfam. Pottioideae > Tortula |
Sibling taxa | ||
Synonyms | Grimmia atrovirens, Desmatodon californicus, Desmatodon convolutus | |
Name authority | Hedwig: Sp. Musc. Frond., 123. (1801) | (Smith) Lindberg: Öfvers. Kongl. Vetensk.-Akad. Förh. 21: 236. (1864) |
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