Tortula mucronifolia |
Tortula modica |
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mucronleaf tortula moss |
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Leaves | oblong-lanceolate to obovate or long-elliptic, apex rounded-acute, mucronate to short-awned, margins recurved at base or somewhat higher, not bordered or bordered at base with 3–5 rows of thicker-walled cells, 1–3:1; costa excurrent, lacking an adaxial pad of cells, distally very gradually narrowing, 3–5 cells across the weakly convex adaxial surface; distal laminal cells rounded-hexagonal, 14–20(–28) µm wide, 1:1, not papillose or very weakly so. |
obovate to spatulate, apex broadly acute or occasionally rounded, short-awned, margins weakly recurved proximally or occasionally plane, weakly bordered distally with 2–4 rows of slightly thicker-walled cells; costa excurrent, lacking an adaxial pad of cells, distally narrow, 2–3 cells across the convex adaxial surface; distal laminal cells irregularly hexagonal or sometimes rectangular, width 20–25(–29) µm wide, 1(–2):1, smooth or 1-papillose (best seen in section). |
Seta | 1–2 cm. |
0.4–0.6 cm. |
Sexual condition | autoicous. |
autoicous. |
Capsule | stegocarpic, not systylius, cylindric, erect and nearly straight, urn 3–6 mm; peristome 1700–2000 µm, teeth of 32 filaments twisted at least 1 full turn, basal membrane high, ca. 1000 µm; operculum ca. 1–2 mm. |
stegocarpic, not systylius, short-cylindric, erect and nearly straight, urn 1–1.5 mm; peristome absent or rudimentary; operculum ca. 0.5 mm. |
Spores | 12–18 µm, spheric, finely papillose. |
30–35 µm, spheric, densely papillose. |
Sporophytes | exerted. |
exerted. |
Tortula mucronifolia |
Tortula modica |
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Phenology | Capsules mature summer. | Capsules mature fall–spring. |
Habitat | Soil, calcareous soil, silt, rock, cliffs, walls | Calcareous soil, old fields, pastures, rocky areas |
Elevation | low to high elevations (0-2700 m) (low to high elevations (0-8900 ft)) | low and moderate elevations |
Distribution |
AK; AZ; CA; CO; CT; IA; ID; IL; IN; MA; MI; MN; MT; ND; NE; NM; NV; NY; OH; OR; PA; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; QC; SK; YT; n Asia; Greenland; Europe; Africa
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NY; PA; ON; Europe; Asia; n Africa |
Discussion | Tortula mucronifolia is quite close to T. subulata, differing largely in the clear medial laminal cells of the former, which are not or rarely papillose, and the marginal border less strongly developed or absent. Where their ranges overlap, intermediates may be found. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
A report of Tortula modica from Mexico was excluded by R. H. Zander (1993). The distribution and characters of this uncommon species were discussed by H. A. Crum and L. E. Anderson (1981), P. M. Eckel (1987), and C. Williams (1966b). Essentially, Tortula modica is a high polyploid (n = 52) taxon distinguished from T. truncata by the larger habit, leaf margins usually recurved proximally, capsules slightly longer, short-cylindric, not tapering to the base, and peristome sometimes present though rudimentary. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 601. | FNA vol. 27, p. 598. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Tortula | Pottiaceae > subfam. Pottioideae > Tortula |
Sibling taxa | ||
Synonyms | Gymnostomum intermedium, Pottia intermedia, Pottia truncata var. major | |
Name authority | Schwägrichen: Sp. Musc. Frond. Suppl. 1(1): 136, plate 34. (1811) | R. H. Zander: Bull. Buffalo Soc. Nat. Sci. 32: 226. (1993) |
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