Tortula acaulon |
Tortula systylia |
|
---|---|---|
cuspidate earth-moss, tooth phascum moss |
desmatodon moss |
|
Leaves | ovate to oblong-lanceolate, apex acute, mucronate to short-awned, occasionally comparatively long-awned, margins recurved proximally to nearly plane, not or weakly bordered distally with slightly thicker-walled cells; costa excurrent, lacking an adaxial pad of cells, distally narrow, 2(–3) cells across adaxial surface; distal laminal cells distally quadrate-hexagonal, width 13–17 µm, 1:1, weakly simply papillose or smooth. |
obovate to occasionally ovate, apex acute or broadly acute, awned, margins recurved or plane proximally, not bordered; costa excurrent into the awn, lacking an adaxial pad of cells, distally narrow, 2–3 cells across the convex adaxial surface; distal laminal cells hexagonal to rhomboidal, width 15–22 µm wide, 1:1 near apex but elongate at mid leaf and basally, smooth. |
Seta | very short. |
0.8–1.1 cm. |
Sexual condition | autoicous. |
autoicous. |
Capsule | cleistocarpic, spheric to very short-ellipsoid, erect, mostly 0.9–1.3 mm; peristome absent; operculum not differentiated. |
systylius, short-cylindric, erect and nearly straight, urn 1.5–2 mm; peristome 200–250 µm, teeth 16, straight, divided to near base but variously fused or perforate, basal membrane low; operculum 0.5–0.7 mm. |
Spores | (25–)33–40 µm, spheric, densely papillose. |
22–30(–35) µm, spheric, warty papillose. |
Sporophytes | immersed. |
exerted. |
Tortula acaulon |
Tortula systylia |
|
Phenology | Capsules mature winter–spring. | Capsules mature summer. |
Habitat | Soil, lawns, fields, banks | Soil in rock crevices, tundra |
Elevation | low to moderate elevations | high elevations (0–3700 m) (high elevations (0–12100 ft)) |
Distribution |
AK; AR; AZ; CA; CO; CT; DE; FL; IA; ID; IL; IN; KS; LA; MD; MI; MN; MO; MT; NC; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; UT; VA; WA; WI; WV; AB; BC; NS; ON; QC; SK; Mexico; Europe; Asia; n Africa; Pacific Islands (New Zealand)
|
AK; CA; CO; WY; AB; BC; MB; NF; NT; NU; QC; YT; Greenland; Europe; Asia |
Discussion | The varieties pilifera and schreberiana are not recognized here. The former name may be used for plants with long awns, and the latter for robust specimens ranging to 1 cm tall, but intergrades appear to be common. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Tortula systylia may be confused with Stegonia but the latter has a distinct triangle of echlorophyllose cells at the apex of the much broader leaves. It is similar also to T. hoppeana, but the laminal cells are smooth. After dehiscence, the systylius capsules retain a raised operculum on the columellar stalk. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 595. | FNA vol. 27, p. 599. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Tortula | Pottiaceae > subfam. Pottioideae > Tortula |
Sibling taxa | ||
Synonyms | Phascum acaulon, Microbryum floerkeanum var. henrici, Phascum cuspidatum, Phascum cuspidatum var. americanum, Phascum cuspidatum var. henrici, Phascum cuspidatum var. piliferum, Phascum cuspidatum var. schreberianum, T. acaulon var. pilifera, T. acaulon var. schreberiana, T. atherodes, T. atherodes var. pilifera, T. atherodes var. schreberiana | Desmatodon systylius |
Name authority | (Withering) R. H. Zander: Bull. Buffalo Soc. Nat. Sci. 32: 378. (1994) | (Schimper) Lindberg: Musc. Scand., 20. (1879) |
Web links |
|
|