Tortula acaulon |
Tortula mucronifolia |
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cuspidate earth-moss, tooth phascum moss |
mucronleaf tortula moss |
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Leaves | ovate to oblong-lanceolate, apex acute, mucronate to short-awned, occasionally comparatively long-awned, margins recurved proximally to nearly plane, not or weakly bordered distally with slightly thicker-walled cells; costa excurrent, lacking an adaxial pad of cells, distally narrow, 2(–3) cells across adaxial surface; distal laminal cells distally quadrate-hexagonal, width 13–17 µm, 1:1, weakly simply papillose or smooth. |
oblong-lanceolate to obovate or long-elliptic, apex rounded-acute, mucronate to short-awned, margins recurved at base or somewhat higher, not bordered or bordered at base with 3–5 rows of thicker-walled cells, 1–3:1; costa excurrent, lacking an adaxial pad of cells, distally very gradually narrowing, 3–5 cells across the weakly convex adaxial surface; distal laminal cells rounded-hexagonal, 14–20(–28) µm wide, 1:1, not papillose or very weakly so. |
Seta | very short. |
1–2 cm. |
Sexual condition | autoicous. |
autoicous. |
Capsule | cleistocarpic, spheric to very short-ellipsoid, erect, mostly 0.9–1.3 mm; peristome absent; operculum not differentiated. |
stegocarpic, not systylius, cylindric, erect and nearly straight, urn 3–6 mm; peristome 1700–2000 µm, teeth of 32 filaments twisted at least 1 full turn, basal membrane high, ca. 1000 µm; operculum ca. 1–2 mm. |
Spores | (25–)33–40 µm, spheric, densely papillose. |
12–18 µm, spheric, finely papillose. |
Sporophytes | immersed. |
exerted. |
Tortula acaulon |
Tortula mucronifolia |
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Phenology | Capsules mature winter–spring. | Capsules mature summer. |
Habitat | Soil, lawns, fields, banks | Soil, calcareous soil, silt, rock, cliffs, walls |
Elevation | low to moderate elevations | low to high elevations (0-2700 m) (low to high elevations (0-8900 ft)) |
Distribution |
AK; AR; AZ; CA; CO; CT; DE; FL; IA; ID; IL; IN; KS; LA; MD; MI; MN; MO; MT; NC; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; UT; VA; WA; WI; WV; AB; BC; NS; ON; QC; SK; Mexico; Europe; Asia; n Africa; Pacific Islands (New Zealand)
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AK; AZ; CA; CO; CT; IA; ID; IL; IN; MA; MI; MN; MT; ND; NE; NM; NV; NY; OH; OR; PA; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NF; NS; NT; NU; ON; QC; SK; YT; n Asia; Greenland; Europe; Africa
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Discussion | The varieties pilifera and schreberiana are not recognized here. The former name may be used for plants with long awns, and the latter for robust specimens ranging to 1 cm tall, but intergrades appear to be common. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Tortula mucronifolia is quite close to T. subulata, differing largely in the clear medial laminal cells of the former, which are not or rarely papillose, and the marginal border less strongly developed or absent. Where their ranges overlap, intermediates may be found. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 595. | FNA vol. 27, p. 601. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Tortula | Pottiaceae > subfam. Pottioideae > Tortula |
Sibling taxa | ||
Synonyms | Phascum acaulon, Microbryum floerkeanum var. henrici, Phascum cuspidatum, Phascum cuspidatum var. americanum, Phascum cuspidatum var. henrici, Phascum cuspidatum var. piliferum, Phascum cuspidatum var. schreberianum, T. acaulon var. pilifera, T. acaulon var. schreberiana, T. atherodes, T. atherodes var. pilifera, T. atherodes var. schreberiana | |
Name authority | (Withering) R. H. Zander: Bull. Buffalo Soc. Nat. Sci. 32: 378. (1994) | Schwägrichen: Sp. Musc. Frond. Suppl. 1(1): 136, plate 34. (1811) |
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