Tortula acaulon |
Tortula inermis |
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cuspidate earth-moss, tooth phascum moss |
tortula moss |
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Leaves | ovate to oblong-lanceolate, apex acute, mucronate to short-awned, occasionally comparatively long-awned, margins recurved proximally to nearly plane, not or weakly bordered distally with slightly thicker-walled cells; costa excurrent, lacking an adaxial pad of cells, distally narrow, 2(–3) cells across adaxial surface; distal laminal cells distally quadrate-hexagonal, width 13–17 µm, 1:1, weakly simply papillose or smooth. |
lingulate, apex rounded-acute, apiculate to short-mucronate, margins recurved from base to near the apex, not bordered; costa subpercurrent, percurrent or very short-excurrent, lacking an adaxial pad of cells but distally narrowing or weakly thickened, 3–4(–5) cells across the convex adaxial surface; distal laminal cells hexagonal, (12–)15–18 µm wide, 1:1, strongly papillose with 4–5 2-fid papillae. |
Seta | very short. |
1.2–1.5(–2.5) cm. |
Sexual condition | autoicous. |
gonioautoicous. |
Capsule | cleistocarpic, spheric to very short-ellipsoid, erect, mostly 0.9–1.3 mm; peristome absent; operculum not differentiated. |
stegocarpic, not systylius, cylindric, erect and nearly straight, urn usually 3–4 mm; peristome length 1000–1500 µm, teeth of 32 filaments twisted at least one full turn, basal membrane 300–600 µm; operculum 1.3–1.6(–2) mm. |
Spores | (25–)33–40 µm, spheric, densely papillose. |
11–15 µm, spheric, finely papillose. |
Sporophytes | immersed. |
exerted. |
Tortula acaulon |
Tortula inermis |
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Phenology | Capsules mature winter–spring. | Capsules mature spring. |
Habitat | Soil, lawns, fields, banks | Soil, rock |
Elevation | low to moderate elevations | low to high elevations (0-2000 m) (low to high elevations (0-6600 ft)) |
Distribution |
AK; AR; AZ; CA; CO; CT; DE; FL; IA; ID; IL; IN; KS; LA; MD; MI; MN; MO; MT; NC; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; UT; VA; WA; WI; WV; AB; BC; NS; ON; QC; SK; Mexico; Europe; Asia; n Africa; Pacific Islands (New Zealand)
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AZ; CA; CO; NM; NV; SD; TX; UT; WY; Mexico (Baja California, Chihuahua, Sonora); Europe; s Asia; n Africa |
Discussion | The varieties pilifera and schreberiana are not recognized here. The former name may be used for plants with long awns, and the latter for robust specimens ranging to 1 cm tall, but intergrades appear to be common. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Tortula inermis has the aspect of a Syntrichia with its ligulate, apiculate leaves and strong costa, but the plant is yellow or orange in KOH solution, and the costal section reveals a rounded stereid band. It is related to T. subulata and T. mucronifolia but the lack of a strong mucro and the narrow but nearly complete recurving of the leaf margins are diagnostic. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 595. | FNA vol. 27, p. 600. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Tortula | Pottiaceae > subfam. Pottioideae > Tortula |
Sibling taxa | ||
Synonyms | Phascum acaulon, Microbryum floerkeanum var. henrici, Phascum cuspidatum, Phascum cuspidatum var. americanum, Phascum cuspidatum var. henrici, Phascum cuspidatum var. piliferum, Phascum cuspidatum var. schreberianum, T. acaulon var. pilifera, T. acaulon var. schreberiana, T. atherodes, T. atherodes var. pilifera, T. atherodes var. schreberiana | Syntrichia subulata var. inermis, Syntrichia inermis |
Name authority | (Withering) R. H. Zander: Bull. Buffalo Soc. Nat. Sci. 32: 378. (1994) | (Bridel) Montagne: Arch. Bot. (Leipzig) 1: 136. (1832) |
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