Tortula acaulon |
Tortula hoppeana |
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cuspidate earth-moss, tooth phascum moss |
wideleaf desmatodon moss |
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Leaves | ovate to oblong-lanceolate, apex acute, mucronate to short-awned, occasionally comparatively long-awned, margins recurved proximally to nearly plane, not or weakly bordered distally with slightly thicker-walled cells; costa excurrent, lacking an adaxial pad of cells, distally narrow, 2(–3) cells across adaxial surface; distal laminal cells distally quadrate-hexagonal, width 13–17 µm, 1:1, weakly simply papillose or smooth. |
ovate to oblong, apex broadly acute or occasionally rounded, apiculate or short- to long-mucronate, occasionally short-awned, margins recurved in middle 2/3 of leaf or occasionally almost plane, not or very weakly bordered distally with ca. 2 rows of rhomboid, less papillose cells; costa subpercurrent, percurrent or excurrent, lacking an adaxial pad of cells, distally narrow, 2–3 cells across the convex adaxial surface; distal laminal cells hexagonal, 15–20 µm wide, 1:1, densely papillose with several 2-fid papillae. |
Seta | very short. |
0.9–1.2(–1.9) cm. |
Sexual condition | autoicous. |
autoicous. |
Capsule | cleistocarpic, spheric to very short-ellipsoid, erect, mostly 0.9–1.3 mm; peristome absent; operculum not differentiated. |
stegocarpic, rarely systylius, cylindric, erect and nearly straight, urn 1.5–2(–2.8) mm; peristome (150–)300–350 length µm, teeth 16, usually not twisted, divided to near the base or perforate basally, the teeth often basally fused in 2s or 4s, basal membrane very low; operculum 0.7–1 mm. |
Spores | (25–)33–40 µm, spheric, densely papillose. |
ca. 20–23(–25) µm, spheric or elliptic, densely papillose. |
Sporophytes | immersed. |
exerted. |
Tortula acaulon |
Tortula hoppeana |
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Phenology | Capsules mature winter–spring. | Capsules mature summer. |
Habitat | Soil, lawns, fields, banks | Soil, calcareous silt |
Elevation | low to moderate elevations | low (in northern regions) to high elevations of over 3300 m (low (in northern regions) to high elevations of over 10800 ft) |
Distribution |
AK; AR; AZ; CA; CO; CT; DE; FL; IA; ID; IL; IN; KS; LA; MD; MI; MN; MO; MT; NC; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; UT; VA; WA; WI; WV; AB; BC; NS; ON; QC; SK; Mexico; Europe; Asia; n Africa; Pacific Islands (New Zealand)
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AK; AZ; CA; CO; ID; MI; MT; NM; NV; OR; UT; WA; WY; AB; BC; NL; NT; NU; QC; YT; n Africa; Greenland; Asia; Europe; Atlantic Islands (Iceland) |
Discussion | The varieties pilifera and schreberiana are not recognized here. The former name may be used for plants with long awns, and the latter for robust specimens ranging to 1 cm tall, but intergrades appear to be common. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Care must be taken to not be confused by recent nomenclatural changes involving the epithet “latifolia” in the Pottioideae. Pottia latifolia is now to be found in Stegonia; Desmatodon latifolius is now Tortula hoppeana; Tortula latifolia is now Syntrichia latifolia. The strong 2-fid laminal papillae of Tortula hoppeana are distinctive, contributing to a crenulate leaf margin at the apex, which is surmounted, excepting specimens with muticous leaves, by a narrow, long mucro of smooth, clear, elongate cells. This is a very common species in subalpine and alpine areas, and in tundra. The present treatment agrees with modern perception (L. E. Anderson et al. 1990; M. J. Cano et al. 2006) that the muticous-leaved variant, which has much the same distribution as the piliferous expression, is not worth recognizing. The peristome teeth are essentially 16 but are commonly paired or in larger groups. Like T. cernua, T. laureri, T. leucostoma, T. obtusifolia, and T. systylia, this species has a capsule mainly maturing in summer, a feature supporting recognition of the group under the name Desmatodon, but possibly at only the subgenus level. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 27, p. 595. | FNA vol. 27, p. 598. |
Parent taxa | Pottiaceae > subfam. Pottioideae > Tortula | Pottiaceae > subfam. Pottioideae > Tortula |
Sibling taxa | ||
Synonyms | Phascum acaulon, Microbryum floerkeanum var. henrici, Phascum cuspidatum, Phascum cuspidatum var. americanum, Phascum cuspidatum var. henrici, Phascum cuspidatum var. piliferum, Phascum cuspidatum var. schreberianum, T. acaulon var. pilifera, T. acaulon var. schreberiana, T. atherodes, T. atherodes var. pilifera, T. atherodes var. schreberiana | Trichostomum hoppeanum, Desmatodon latifolius, Desmatodon latifolius var. muticus, T. eucalyptrata, T. euryphylla |
Name authority | (Withering) R. H. Zander: Bull. Buffalo Soc. Nat. Sci. 32: 378. (1994) | (Schultz) Ochyra: Bryologist 107: 499. (2005) |
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