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cuspidate earth-moss, tooth phascum moss

Habit Plants green to tan.
Stem

central strand usually present, sclerodermis and hyalodermis commonly absent.

Leaves

ovate to oblong-lanceolate, apex acute, mucronate to short-awned, occasionally comparatively long-awned, margins recurved proximally to nearly plane, not or weakly bordered distally with slightly thicker-walled cells;

costa excurrent, lacking an adaxial pad of cells, distally narrow, 2(–3) cells across adaxial surface;

distal laminal cells distally quadrate-hexagonal, width 13–17 µm, 1:1, weakly simply papillose or smooth.

usually ovate or broadly ligulate to spathulate, apex usually rounded or broadly acute, margins recurved basally, distal laminal cells if 2-stratose stacked directly over one another, walls smooth or papillose, KOH yellow or red, occasionally orange, occasionally yellow distally and red basally;

costa with or without a differentiated abaxial epidermis;

gemmae largely absent.

Seta

very short.

Sexual condition

autoicous.

Capsule

cleistocarpic, spheric to very short-ellipsoid, erect, mostly 0.9–1.3 mm;

peristome absent;

operculum not differentiated.

Spores

(25–)33–40 µm, spheric, densely papillose.

Sporophytes

immersed.

Tortula acaulon

Pottiaceae subfam. pottioideae

Phenology Capsules mature winter–spring.
Habitat Soil, lawns, fields, banks
Elevation low to moderate elevations
Distribution
from FNA
AK; AR; AZ; CA; CO; CT; DE; FL; IA; ID; IL; IN; KS; LA; MD; MI; MN; MO; MT; NC; NE; NJ; NM; NV; NY; OH; OK; OR; PA; SD; TN; TX; UT; VA; WA; WI; WV; AB; BC; NS; ON; QC; SK; Mexico; Europe; Asia; n Africa; Pacific Islands (New Zealand)
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Worldwide
Discussion

The varieties pilifera and schreberiana are not recognized here. The former name may be used for plants with long awns, and the latter for robust specimens ranging to 1 cm tall, but intergrades appear to be common.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 21 (15 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 595. FNA vol. 27, p. 585.
Parent taxa Pottiaceae > subfam. Pottioideae > Tortula Pottiaceae
Sibling taxa
T. amplexa, T. atrovirens, T. bolanderi, T. brevipes, T. californica, T. cernua, T. cuneifolia, T. deciduidentata, T. guepinii, T. hoppeana, T. inermis, T. lanceola, T. laureri, T. leucostoma, T. modica, T. mucronifolia, T. muralis, T. nevadensis, T. obtusifolia, T. plinthobia, T. porteri, T. protobryoides, T. subulata, T. systylia, T. truncata
Subordinate taxa
Synonyms Phascum acaulon, Microbryum floerkeanum var. henrici, Phascum cuspidatum, Phascum cuspidatum var. americanum, Phascum cuspidatum var. henrici, Phascum cuspidatum var. piliferum, Phascum cuspidatum var. schreberianum, T. acaulon var. pilifera, T. acaulon var. schreberiana, T. atherodes, T. atherodes var. pilifera, T. atherodes var. schreberiana
Name authority (Withering) R. H. Zander: Bull. Buffalo Soc. Nat. Sci. 32: 378. (1994) Brotherus: in H. G. A. Engler et al., Nat. Pflanzenfam. ed. 2, 10: 282. (1924)
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