Tonestus pygmaeus |
Tonestus eximius |
|
---|---|---|
pygmy goldenweed, pygmy serpentweed |
Lake Tahoe serpentweed, Tahoe tonestus |
|
Habit | Perennials, loosely cespitose, 1–9 cm; with branched caudices surmounting well-developed taproots. | Perennials, cespitose, 5–13 cm; with branched caudices surmounting ± well-developed taproots, or short rhizomes. |
Stems | densely hairy, sometimes also stipitate-glandular. |
(clothed with marcescent leaf bases) glabrous, densely stipitate-glandular. |
Leaves | basal blades linear to spatulate, 10–50(–95) × 1.5–6 mm; cauline blades oblong to broadly lanceolate, 15–34 × 3–7 mm, gradually reduced distally; 3–5-nerved, margins entire, ciliate, sometimes stipitate-glandular, faces eglandular or sparsely stipitate-glandular. |
basal blades broadly spatulate, 20–35 × 12–15 mm, margins coarsely, irregularly serrate to dentate; cauline blades oblong, 9–52 × 5–15 mm, ± reduced distally, margins coarsely, irregularly serrate or nearly entire; 1- or 3-nerved, margins stipitate-glandular, faces glabrous, stipitate-glandular. |
Peduncles | 3–8 mm. |
2–15 mm. |
Involucres | broadly campanulate, 8–20 × 7–12 mm. |
broadly campanulate, 9–16(–20) × 5–11(–15) mm. |
Ray florets | 10–16(–35); laminae elliptic, 6.5–8.5 × 1.5–3.5 mm. |
10–13; laminae elliptic, 5–8 × 1–3 mm. |
Disc florets | 43–66; corollas narrowly funnelform, 4.5–7.5 mm, lobes erect to spreading, 0.8–1.2 mm, lengths ca. 1/7 corollas; anthers 2.2–2.8 mm; style-branch appendages lanceolate, 0.8–1.1 × 0.3 mm wide, stigmatic lines 0.6–1 mm. |
41–49; corollas narrowly funnelform, 5.5–7.5(–8.2) mm, exceeding involucres, lobes erect, 0.7–1.7 mm, 1/10–1/5 corolla length; anthers 2.5–3.3 mm; style-branch appendages subulate, 0.9–1.3 mm, stigmatic lines 0.9–1.6 mm. |
Phyllaries | 16–34, in 3–4 (–5) series, 1- or weakly 3–5-nerved, ± equal, margins ciliolate at least distally; outer and mid green, ovate to oblong, 4.5–10.5 × 1.5–5.5 mm, foliaceous, apices rounded to obtuse, faces sparsely stipitate-glandular distally; inner green or usually anthocyanic, linear to narrowly oblong, 5–9 × 0.9–4 mm, chartaceous proximally, margins scarious, often fimbriate, sometimes stipitate-glandular, apices acute to acuminate, sometimes reflexed, faces sparsely stipitate-glandular distally. |
16–32 in 3–4 series, 1- or weakly 3-nerved, ± equal, margins eciliate; outer and mid green, oblong, 6–10 × 2–5 mm, foliaceous, apices rounded or obtuse to acute; inner green or sometimes anthocyanic, linear, 7–9 × 0.5–2 mm, chartaceous proximally, margins scarious, fimbriate and/or stipitate-glandular distally, apices acute to acuminate, often reflexed, faces stipitate-glandular distally. |
Heads | 1(–2). |
usually borne singly, sometimes 2–4. |
Cypselae | cylindric, 2–5 mm, 8–9-nerved, faces villous; pappus bristles ± 34–51, flexible. |
linear, 2.5–6 mm, 5–9-nerved, faces villous, often ± viscid; pappus bristles 19–43, brittle. |
2n | = 18. |
= 18. |
Tonestus pygmaeus |
Tonestus eximius |
|
Phenology | Flowering and fruiting early–late summer. | Flowering and fruiting early–late summer. |
Habitat | Meadows, fellfields, talus slopes, rock crevices, dwarf coniferous forests in alpine and upper subalpine communities, on soils of usually granitic, sometimes limestone origin | Crevices of granitic outcrops, talus slopes, in alpine and coniferous subalpine communities |
Elevation | 3100–4100 m (10200–13500 ft) | 1800–3300 m (5900–10800 ft) |
Distribution |
CO; MT; NM; WY
|
CA; NV |
Discussion | Tonestus pygmaeus is uncommon in Wyoming, known only from the Snowy Range of the Medicine Bow Mountains. Attribution of this species to Montana is based on historic collections. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Tonestus eximius is known only from the upper elevations of the Sierra Nevada in the vicinity of Lake Tahoe in Alpine and El Dorado counties, California, and Washoe County, Nevada. It is similar to Lorandersonia peirsonii, although smaller in most respects. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 183. | FNA vol. 20, p. 183. |
Parent taxa | Asteraceae > tribe Astereae > Tonestus | Asteraceae > tribe Astereae > Tonestus |
Sibling taxa | ||
Synonyms | Stenotus pygmaeus, Haplopappus pygmaeus | Haplopappus eximius |
Name authority | (Torrey & A. Gray) A. Nelson: Bot. Gaz. 37: 262. (1904) | (H. M. Hall) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 70. (1918) |
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