Tonestus lyallii |
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Lyall's goldenweed, Lyall's serpentweed, Lyall's tonenstus, Lyall's tonestus |
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Habit | Perennials, loosely cespitose, 4–8.5(–15) cm; with branched caudices surmounting poorly developed taproots, or short rhizomes. |
Stems | glabrous, densely stipitate-glandular. |
Leaves | basal blades linear to spatulate, 30–85 × 3–12 mm; cauline blades oblong to broadly lanceolate, 12–37 × 3–9(–12) mm, gradually reduced distally; 3–5-nerved, margins entire, eciliate, stipitate-glandular, faces stipitate-glandular. |
Peduncles | 2–5 mm. |
Involucres | broadly campanulate, 11–22 × 8–11 mm. |
Ray florets | 11–23; laminae elliptic, 6.5–8 × 1–3.5 mm. |
Disc florets | 27–56; corollas narrowly funnelform, 5.5–8.5 mm, scarcely exceeding involucres, lobes erect to spreading, 0.6–0.8 mm, lengths 1/10–1/5 corollas; anthers 2–2.5 mm; style-branch appendages lanceolate, 0.7–1 × 0.2–0.3 mm, stigmatic lines 0.9–1.4 mm. |
Phyllaries | 18–42, in 3–4(–5) series, , 1- or weakly 3-nerved, ± equal, margins eciliate; outer and mid oblong to broadly lanceolate, 5.2–9.5 × 1–3.5 mm, stipitate-glandular, apices acute; inner green or anthocyanic, linear, 8–13 × 1–2 mm, chartaceous proximally, margins scarious, fimbriate and/or stipitate-glandular distally, apices acute to acuminate, sometimes reflexed, faces stipitate-glandular distally. |
Heads | 1(–2). |
Cypselae | linear, 2.5–5.5 mm, 8–12-nerved, faces glabrous or sparsely hairy distally; pappus bristles 35–52, flexible. |
2n | = 18. |
Tonestus lyallii |
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Phenology | Flowering and fruiting early–late summer. |
Habitat | Meadows, fellfields, talus slopes, rock crevices, open coniferous forests in alpine and subalpine communities, on soils usually of granitic, sometime of limestone or shale origin |
Elevation | 1500–3800 m (4900–12500 ft) |
Distribution |
CA; CO; ID; MT; NV; OR; WA; WY; BC; Alta
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Discussion | Tonestus lyallii is widespread in the central Rocky Mountains and ranges of the Pacific Northwest, and is known in the Great Basin from collections in the Ruby Mountains in Elko County, Nevada. Populations documented from the Coast Range in Siskyou and Trinity counties, California, are disjunct from those in Oregon and Washington by more than 700 km. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 183. |
Parent taxa | Asteraceae > tribe Astereae > Tonestus |
Sibling taxa | |
Synonyms | Haplopappus lyallii |
Name authority | (A. Gray) A. Nelson: Bot. Gaz. 37: 262. (1904) |
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