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Lyall's goldenweed, Lyall's serpentweed, Lyall's tonenstus, Lyall's tonestus

Habit Perennials, loosely cespitose, 4–8.5(–15) cm; with branched caudices surmounting poorly developed taproots, or short rhizomes.
Stems

glabrous, densely stipitate-glandular.

Leaves

basal blades linear to spatulate, 30–85 × 3–12 mm;

cauline blades oblong to broadly lanceolate, 12–37 × 3–9(–12) mm, gradually reduced distally; 3–5-nerved, margins entire, eciliate, stipitate-glandular, faces stipitate-glandular.

Peduncles

2–5 mm.

Involucres

broadly campanulate, 11–22 × 8–11 mm.

Ray florets

11–23;

laminae elliptic, 6.5–8 × 1–3.5 mm.

Disc florets

27–56;

corollas narrowly funnelform, 5.5–8.5 mm, scarcely exceeding involucres, lobes erect to spreading, 0.6–0.8 mm, lengths 1/10–1/5 corollas;

anthers 2–2.5 mm;

style-branch appendages lanceolate, 0.7–1 × 0.2–0.3 mm, stigmatic lines 0.9–1.4 mm.

Phyllaries

18–42, in 3–4(–5) series, , 1- or weakly 3-nerved, ± equal, margins eciliate;

outer and mid oblong to broadly lanceolate, 5.2–9.5 × 1–3.5 mm, stipitate-glandular, apices acute;

inner green or anthocyanic, linear, 8–13 × 1–2 mm, chartaceous proximally, margins scarious, fimbriate and/or stipitate-glandular distally, apices acute to acuminate, sometimes reflexed, faces stipitate-glandular distally.

Heads

1(–2).

Cypselae

linear, 2.5–5.5 mm, 8–12-nerved, faces glabrous or sparsely hairy distally;

pappus bristles 35–52, flexible.

2n

= 18.

Tonestus lyallii

Phenology Flowering and fruiting early–late summer.
Habitat Meadows, fellfields, talus slopes, rock crevices, open coniferous forests in alpine and subalpine communities, on soils usually of granitic, sometime of limestone or shale origin
Elevation 1500–3800 m (4900–12500 ft)
Distribution
from FNA
CA; CO; ID; MT; NV; OR; WA; WY; BC; Alta
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[BONAP county map]
Discussion

Tonestus lyallii is widespread in the central Rocky Mountains and ranges of the Pacific Northwest, and is known in the Great Basin from collections in the Ruby Mountains in Elko County, Nevada. Populations documented from the Coast Range in Siskyou and Trinity counties, California, are disjunct from those in Oregon and Washington by more than 700 km.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 20, p. 183.
Parent taxa Asteraceae > tribe Astereae > Tonestus
Sibling taxa
T. eximius, T. graniticus, T. pygmaeus
Synonyms Haplopappus lyallii
Name authority (A. Gray) A. Nelson: Bot. Gaz. 37: 262. (1904)
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