Timmia austriaca |
Timmia |
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Austria timmia moss, false-polytrichum |
timmia moss |
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Habit | Plants without deciduous distal leaves. | Plants (1–)2–9(–13) cm, in compact or loose tufts, brownish proximally, bright to yellowish green distally. | ||||||||||||
Leaves | apex acute (but with terminal 4–5 cells often truncated); costa subpercurrent; sheath orange but appearing red when still on stem; limbs green, pellucid; limb-sheath transition abrupt, sharply angled; limb margins strongly dentate in distal 1/3–1/2, less dentate to entire proximally, occasionally crenulate or entire throughout; limb lamina cells 8–14(–16) × 8–12(–13) µm, with low conical or rounded mamillae on the adaxial surface, abaxial surface smooth; distal sheath lamina cells smooth or with 1–6(–8) large round, often verrucose papillae over the abaxial surfaces of lumen; cells at leaf insertion not hyaline and fragile; abaxial surface of costa limb smooth, toothed or papillose near apex, adaxial cells with conical or rounded mamillae. |
crisped to imbricate (and occasionally inrolled) when dry, erect-patent to spreading when moist, (1.5–)2–8(–12) mm, 0.5–1.2 mm wide at mid-limb; cells of the sheath lamina longer and narrower near the margins, shorter and broader near the limb-sheath transition, smooth on the adaxial surface, smooth or with a series of linearly-arranged, large round papillae on the abaxial surface of the lumens. |
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Sexual condition | dioicous; perichaetial leaves more strongly dentate than vegetative leaves, sheath up to half the leaf length. |
monoicous or dioicous; male gametophyte (in dioicous taxa) monopodial, with the stem continuing growth through terminal perigonia; paraphyses few, of 10–15(–40) cells, filiform, with fusiform tips; archegonia 1–6 per perichaetium, ca. 1–2 mm; antheridia 5–40 per perigonium, ca. 1–2 mm. |
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Capsule | spirally plicate (even when young and operculate); exothecial cell walls sinuose; stomata mostly on the neck; endostome cilia without appendiculations on the interior surfaces. |
ca. 2.5–3 × 1–1.5 mm; exothecial cells with smooth or sinuose walls; peristome diplolepideous; exostome teeth 0.5–1 mm, yellow in proximal half, pale or hyaline distally; endostome hyaline or yellowish, with a high basal membrane about half the height of the exostome, and 64 irregularly anastomosing, nodose cilia papillose externally and smooth or appendiculate on the interior surface. |
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Calyptra | without longitudinal split, not persistent on seta. |
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Spores | round, finely papillose, pale yellow or brown, 13–22 µm. |
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Timmia austriaca |
Timmia |
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Phenology | Sporophytes sporadic throughout the range. | |||||||||||||
Habitat | but more frequent in non-Arctic regions, various habitats including dry exposed ridges, wet river edges or forested valleys | |||||||||||||
Elevation | low to high elevations (0-3000 m) (low to high elevations (0-9800 ft)) | |||||||||||||
Distribution |
AK; CA; CO; ID; MI; MT; NM; NV; OR; SD; WA; WY; AB; BC; NB; NF; NT; NU; QC; SK; YT; Atlantic Islands (Iceland); Greenland; Eurasia (Alps, Greece, Italy, Pyrenees, Scandinavia, United Kingdom); e Asia (Russia)
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North America; Eurasia; n Africa; Pacific Islands (Hawaii, New Zealand) in arctic; boreal; montane; and temperate areas |
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Discussion | A polymorphic yet remarkably well-defined species, Timmia austriaca has the most well-delimited sheath of all Timmia taxa, with an abrupt change in color and sharp angle at the limb-sheath transition, and it is the only taxon where the costa widens slightly just at the limb-sheath transition. Its distinct habit and leaf angularity, dioicous sexuality, non-appendiculate endostome cilia, and orange sheaths separate it from T. megapolitana, while its habit, leaf angularity, and non-deciduous leaves will separate it from T. norvegica and T. sibirica. The perigonial leaves are short and broad, consisting mostly of sheath. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 4 (4 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 27. | FNA vol. 27, p. 165. | ||||||||||||
Parent taxa | Timmiaceae > Timmia | Timmiaceae | ||||||||||||
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Subordinate taxa | ||||||||||||||
Synonyms | T. arctica, T. austriaca var. arctica, T. austriaca var. brevifolia, T. austriaca var. papillosa | |||||||||||||
Name authority | Hedwig: Sp. Musc. Frond., 176, plate 42, figs. 1–7. (1801) | Hedwig: Sp. Musc. Frond., 176. 1801 , | ||||||||||||
Web links |