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Austria timmia moss, false-polytrichum

Habit Plants without deciduous distal leaves.
Leaf

apex acute (but with terminal 4–5 cells often truncated);

costa subpercurrent;

sheath orange but appearing red when still on stem;

limbs green, pellucid;

limb-sheath transition abrupt, sharply angled;

limb margins strongly dentate in distal 1/3–1/2, less dentate to entire proximally, occasionally crenulate or entire throughout;

limb lamina cells 8–14(–16) × 8–12(–13) µm, with low conical or rounded mamillae on the adaxial surface, abaxial surface smooth;

distal sheath lamina cells smooth or with 1–6(–8) large round, often verrucose papillae over the abaxial surfaces of lumen;

cells at leaf insertion not hyaline and fragile;

abaxial surface of costa limb smooth, toothed or papillose near apex, adaxial cells with conical or rounded mamillae.

Sexual condition

dioicous;

perichaetial leaves more strongly dentate than vegetative leaves, sheath up to half the leaf length.

Capsule

spirally plicate (even when young and operculate);

exothecial cell walls sinuose;

stomata mostly on the neck;

endostome cilia without appendiculations on the interior surfaces.

Calyptra

without longitudinal split, not persistent on seta.

Timmia austriaca

Phenology Sporophytes sporadic throughout the range.
Habitat but more frequent in non-Arctic regions, various habitats including dry exposed ridges, wet river edges or forested valleys
Elevation low to high elevations (0-3000 m) (low to high elevations (0-9800 ft))
Distribution
from FNA
AK; CA; CO; ID; MI; MT; NM; NV; OR; SD; WA; WY; AB; BC; NB; NF; NT; NU; QC; SK; YT; Atlantic Islands (Iceland); Greenland; Eurasia (Alps, Greece, Italy, Pyrenees, Scandinavia, United Kingdom); e Asia (Russia)
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Discussion

A polymorphic yet remarkably well-defined species, Timmia austriaca has the most well-delimited sheath of all Timmia taxa, with an abrupt change in color and sharp angle at the limb-sheath transition, and it is the only taxon where the costa widens slightly just at the limb-sheath transition. Its distinct habit and leaf angularity, dioicous sexuality, non-appendiculate endostome cilia, and orange sheaths separate it from T. megapolitana, while its habit, leaf angularity, and non-deciduous leaves will separate it from T. norvegica and T. sibirica. The perigonial leaves are short and broad, consisting mostly of sheath.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27.
Parent taxa Timmiaceae > Timmia
Sibling taxa
T. megapolitana, T. norvegica, T. sibirica
Synonyms T. arctica, T. austriaca var. arctica, T. austriaca var. brevifolia, T. austriaca var. papillosa
Name authority Hedwig: Sp. Musc. Frond., 176, plate 42, figs. 1–7. (1801)
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