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five-needle pricklyleaf

pricklyleaf

Habit Perennials or subshrubs, ± grayish to green, to 15(–25) cm, usually puberulent to canescent, sometimes glabrescent or glabrous. Annuals, perennials, subshrubs, or shrubs, (1–)5–30 cm.
Stems

erect or spreading.

erect to spreading or decumbent, branched from bases or ± throughout.

Leaves

mostly opposite;

blades mostly pinnately lobed, 6–28+ mm overall, lobes 3–11 linear to filiform (usually stiff, setiform).

cauline; opposite or alternate; petiolate or sessile;

blades often pinnately lobed, blades or lobes spatulate to linear or filiform, ultimate margins entire or toothed, faces glabrous or arachnose, canescent, floccose, lanate, puberulent, or tomentose (bases little, if at all, bristly-ciliate, oil-glands scattered in laminae or submarginal).

Peduncles

20–100 mm, puberulent or glabrous.

Involucres

obconic to campanulate or hemispheric, 4–6 mm.

campanulate to obconic, 2–7 mm diam.

Receptacles

convex, smooth or ± pitted, epaleate.

Ray florets

(8–)12–21;

corollas yellow to orange-yellow, laminae 2–6(–8) × 1–3 mm.

usually 5, 8, 13, or 21, pistillate, fertile;

corollas usually yellow to orange, rarely white.

Disc florets

16–40 or 50–80;

corollas yellow, 2–4 mm (tending to zygomorphy in peripheral florets in some plants).

16–100+, bisexual, fertile;

corollas yellow to orange, tubes shorter than ± cylindric throats, lobes 5, deltate.

Phyllaries

12–21, margins of outer distinct 1/5 to nearly all their lengths, abaxial faces puberulent or glabrous.

persistent, 8–13(–22) in ± 2 series (strongly connate 2/3–7/8+ their lengths, seldom with outer margins distinct more than 1/2 their lengths, exceptions: T. aurea, T. pentachaeta var. belenidium), usually bearing oil-glands.

Calyculi

0, or of 1–5 deltate bractlets, lengths less than 1/2 phyllaries.

0 or of 1–8 deltate to linear bractlets (bearing oil-glands).

Heads

usually radiate, sometimes discoid or radiant, borne singly.

Cypselae

2–3 mm;

pappi of 10 erose and/or aristate scales mostly 1–3 mm.

obpyramidal, obconic, or cylindro-clavate, glabrous or sparsely strigillose;

pappi persistent, coroniform or of 10(–20) distinct scales (each scale erose, or 1–5-aristate, or a fascicle of 5–9 basally connate bristles).

x

= 8.

Thymophylla pentachaeta

Thymophylla

Distribution
from FNA
AZ; CA; NM; NV; TX; UT; Mexico [Introduced(?) in South America (Argentina)]
[WildflowerSearch map]
[BONAP county map]
from USDA
sw United States; Mexico [Introduced in West Indies, South America, Asia, Africa]
[BONAP county map]
Discussion

Varieties 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 13 (8 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucres cylindric, 2–3.5 mm diam.; disc florets 16–40
var. hartwegii
1. Involucres obconic, campanulate, or hemispheric, mostly 4–5 mm diam.; disc florets 50–80
→ 2
2. Peduncles 20–50 mm; margins of outer phyllaries distinct almost to bases
var. belenidium
2. Peduncles (40–)50–100 mm; margins of outer phyllaries distinct less ca. 1/2 or ca. 1/3 their lengths
→ 3
3. Leaf lobes mostly 9–11; phyllaries usually glabrous or glabrescent, rarely hairy
var. pentachaeta
3. Leaf lobes mostly 5–7(–10); phyllaries densely puberulent
var. puberula
1. Leaf blades not lobed (margins entire, toothed, or distally 3-fid); calyculi: lengths of bractlets usually 1/2+ phyllaries
→ 2
1. Leaf blades usually lobed (lobes 3–15+, linear to filiform, blades linear, usually entire in T. tenuiloba var. wrightii); calyculi 0, or lengths of bractlets less than 1/2 phyllaries
→ 4
2. Plants green, usually puberulent, sometimes glabrescent, rarely glabrous; leaves mostly opposite
T. acerosa
2. Plants ashy white, arachnose, lanate, floccose, or tomentose; leaves mostly alternate
→ 3
3. Stems spreading, often prostrate; leaf blades spatulate
T. micropoides
3. Stems erect; leaf blades linear to filiform, sometimes distally 3-fid
T. tephroleuca
4. Perennials or subshrubs (sometimes flowering first year); leaves mostly opposite
→ 5
4. Annuals (rarely persisting); leaves mostly alternate (sometimes opposite at 1–3 proximal nodes)
→ 6
5. Plants green, usually puberulent to canescent, sometimes glabrescent or glabrous; calyculi 0 or of 1–5 deltate bractlets
T. pentachaeta
5. Plants ashy white, tomentose; calyculi of 1–3 subulate bractlets
T. setifolia
6. Calyculi of 3–8 bractlets; disc florets 50–100+
T. tenuiloba
6. Calyculi 0, or of 1–2 bractlets; disc florets 25–45
→ 7
7. Margins of outer phyllaries distinct nearly to bases
T. aurea
7. Margins of outer phyllaries distinct less than 1/2+ their lengths
T. concinna
Source FNA vol. 21, p. 241. FNA vol. 21, p. 239. Author: John L. Strother.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Pectidinae > Thymophylla Asteraceae > tribe Heliantheae > subtribe Pectidinae
Sibling taxa
T. acerosa, T. aurea, T. concinna, T. micropoides, T. setifolia, T. tenuiloba, T. tephroleuca
Subordinate taxa
T. pentachaeta var. belenidium, T. pentachaeta var. hartwegii, T. pentachaeta var. pentachaeta, T. pentachaeta var. puberula
T. acerosa, T. aurea, T. concinna, T. micropoides, T. pentachaeta, T. setifolia, T. tenuiloba, T. tephroleuca
Synonyms Hymenatherum pentachaetum, Dyssodia pentachaeta Dyssodia section Gnaphalopsis, Dyssodia section Hymenatherum, Hymenatherum
Name authority (de Candolle) Small: Fl. S.E. U.S., 1295, 1341. (1903) Lagasca: Gen. Sp. Pl., 25. (1816)
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