Thelypteris simulata |
Thelypteridaceae |
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bog fern, Massachusetts fern, thélyptère simulatrice |
marsh fern family, thelypteris family |
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Habit | Plants terrestrial or on rock [epiphytic]. | |||||||||
Stems | long-creeping, 1.5–3 mm diam. |
creeping to erect, scaly at apex. |
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Leaves | monomorphic, dying back in winter, often 1 cm or more apart, fertile leaves often somewhat longer than sterile leaves, 25–80 cm. |
monomorphic or somewhat dimorphic [dimorphic]. |
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Petiole | straw-colored above base, 12–45 cm × 1–3 mm, at base sparsely set with tan, ovate, glabrous scales. |
in cross section with 2 crescent-shaped vascular bundles at base. |
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Blade | lanceolate, 14–35 cm, proximal pinnae largest, or lowest pair slightly smaller, blade tapered gradually to pinnatifid apex. |
pinnate to pinnate-pinnatifid, rarely more than 2-pinnate [simple]; rachis grooved adaxially or not, grooves not continuous with grooves of next order. |
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Pinnae | deeply pinnatifid to ca. 1 mm from costa, 3–10 × 1–2 cm; segments oblong, somewhat oblique, entire; proximal pair of veins from adjacent segments meeting margin above sinus. |
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Veins | free or anastomosing, running to margin, areoles with or without included free veinlets. |
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Sori | round, medial; indusia tan, often glandular; sporangia glabrous. |
inframedial to supramedial, occasionally nearly marginal, round or oblong, rarely elongate along veins; indusia reniform or sometimes absent. |
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Spores | bilateral, monolete [rarely globose-tetrahedral and trilete], usually with a prominent, crested, echinate, or reticulate perispore. |
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Gametophytes | green, cordate, usually hairy or glandular; antheridia 3-celled. |
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Indument | abaxially of sparsely set hairs 0.2–0.4 mm primarily on costae, also of yellowish short-stalked glands less than 0.1 mm, blade tissue with numerous reddish to orangish, resinous, shiny, sessile, hemispheric glands; blades adaxially with scattered hairs on veins. |
of transparent, needlelike, hooked, septate, or stellate hairs, or rarely hairs lacking. |
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2n | = 128. |
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Thelypteris simulata |
Thelypteridaceae |
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Habitat | Terrestrial in acid soils of shaded swamps and bogs, frequently associated with sphagnum | |||||||||
Elevation | 0–100 m (0–300 ft) | |||||||||
Distribution |
CT; DE; MA; MD; ME; NH; NJ; NY; PA; RI; VA; VT; WI; WV; NB; NS; ON; QC
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Mostly tropical |
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Discussion | Members of Thelypteridaceae have historically been associated with Dryopteridaceae (in particular, Dryopteris) but in fact have no close relationship with that family. Thelypteris and allies differ from Dryopteris and allies by their indument of transparent needlelike hairs (versus needlelike hairs absent in Dryopteridaceae); general absence of blade scales (versus blade scales often present); petiole vasculature in cross section with two crescent-shaped bundles (versus many round bundles arranged in an arc, Athyrium and allies exceptional); generally 1-pinnate to pinnate-pinnatifid blades (versus often more divided); veins usually not forking in the ultimate segments (versus often forking); adaxial grooves discontinuous from rachis to costae, or grooves lacking (versus grooves often continuous); and chromosome base numbers from 27–36 (versus generally 40, 41). Genera 1 to ca. 30, depending on circumscription, species ca. 900 (as circumscribed here, 3 genera and 25 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2, p. 206. | ||||||||
Parent taxa | Thelypteridaceae > Thelypteris > subg. Parathelypteris | |||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Aspidium simulatum, Parathelypteris simulata | |||||||||
Name authority | (Davenport) Nieuwland: Amer. Midl. Naturalist 1: 226. (1910) | Ching ex Pichi Sermolli | ||||||||
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