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fougère de New York, New York fern, thélyptéride de New York

free-tip star-hair fern, freetip maiden fern

Stems

usually long-creeping, 1.5–2.5 mm diam.

short-creeping, 3–5 mm diam.

Leaves

monomorphic, dying back in winter, mostly evenly spaced 1 cm or more (sun-gathering leaves in loose cluster), (25–)40–85 cm.

somewhat dimorphic, evergreen, somewhat spaced, fertile leaves long-petioled, more erect, and with more contracted pinnae, to ca. 1.1 m. Petiole straw-colored, to 60 cm × 2–5 mm, at base sparsely set with brown, lanceolate, stellate-hairy scales.

Petiole

straw-colored, 4–25 cm × 1–3 mm, at base with scales tan to reddish brown, ovate, glabrous.

Blade

elliptic, 15–60 cm, proximal 4–10 pinna pairs gradually smaller toward base (smallest often less than 5 mm), blade tapering gradually to pinnatifid apex.

30–45(–55) × 15–25 cm, broadest at base, with apical pinna similar to lateral pinnae.

Pinnae

deeply pinnatifid to within 1 mm of costa, 3–9(–13) × 1–2(–2.5) cm;

segments oblong to linear, somewhat oblique, entire to crenulate;

proximal pair of veins from adjacent segments meeting margin above sinus.

6–8(–12) pairs, 7–15(–18) × 2–3 cm (fertile 1–2 cm), incised 1/2–3/4 of width;

segments 3–5 mm wide, rounded at apex;

veins 6–10 pairs, proximal pair from adjacent segments united at obtuse angle below sinus with excurrent vein ca. 0.5–1 mm to sinus.

Sori

round, supramedial;

indusia tan, often ciliate;

sporangia glabrous.

round, medial to supramedial;

indusia lacking;

sporangia with numerous hairs 0.1 mm.

Indument

abaxially of moderately to densely set hairs to 1 mm on rachises, costae, and veins, glands lacking or yellowish to light orangish, mostly sessile on blade tissue;

blades adaxially often with hairs on veins.

abaxially of mostly needlelike hairs 0.1–0.3 mm on costae and veins, blade tissue glabrous on both sides.

2n

= 54.

= 144.

Thelypteris noveboracensis

Thelypteris tetragona

Habitat Terrestrial in moist woods, especially near swamps, streams, and in vernal seeps of ravines, often in slightly disturbed secondary forests, frequently forming large colonies Damp woods
Elevation 0–1100 m (0–3600 ft) 0–50 m (0–200 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IL; IN; KY; LA; MA; MD; ME; MI; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WV; NB; NF; NS; ON; PE; QC; SPM
[WildflowerSearch map]
from FNA
FL; Mexico; West Indies in the Antilles; Central America to Panama; n South America
[BONAP county map]
Source FNA vol. 2. FNA vol. 2.
Parent taxa Thelypteridaceae > Thelypteris > subg. Parathelypteris Thelypteridaceae > Thelypteris > subg. Goniopteris
Sibling taxa
T. augescens, T. dentata, T. grandis, T. hispidula, T. interrupta, T. kunthii, T. nevadensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. sclerophylla, T. serrata, T. simulata, T. tetragona
T. augescens, T. dentata, T. grandis, T. hispidula, T. interrupta, T. kunthii, T. nevadensis, T. noveboracensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. sclerophylla, T. serrata, T. simulata
Synonyms Polypodium noveboracense, Dryopteris noveboracensis, Parathelypteris noveboracensis, T. thelypterioides Polypodium tetragonum, Dryopteris tetragona, Goniopteris tetragona
Name authority (Linnaeus) Nieuwland: Amer. Midl. Naturalist 1: 225. (1910) (Swartz) Small: Ferns S. E. States 256. (1938)
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