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fougère de New York, New York fern, thélyptéride de New York

stiff maiden fern, stiff star-hair fern

Stems

usually long-creeping, 1.5–2.5 mm diam.

short-creeping to suberect, 5–8 mm diam.

Leaves

monomorphic, dying back in winter, mostly evenly spaced 1 cm or more (sun-gathering leaves in loose cluster), (25–)40–85 cm.

monomorphic, evergreen, clustered, 20–55(–80) cm.

Petiole

straw-colored, 4–25 cm × 1–3 mm, at base with scales tan to reddish brown, ovate, glabrous.

straw-colored, 5–18(–25) cm × 1–3 mm, at base with brown, lanceolate, stellate-hairy scales.

Blade

elliptic, 15–60 cm, proximal 4–10 pinna pairs gradually smaller toward base (smallest often less than 5 mm), blade tapering gradually to pinnatifid apex.

15–40(–55) × 5–10 cm, proximal 2–5(–10) pairs of pinnae gradually reduced, blade gradually narrowed distally to a pinnatifid apex.

Pinnae

deeply pinnatifid to within 1 mm of costa, 3–9(–13) × 1–2(–2.5) cm;

segments oblong to linear, somewhat oblique, entire to crenulate;

proximal pair of veins from adjacent segments meeting margin above sinus.

18–25 pairs, 2–5(–8) × 1–1.5(–2) cm, deeply serrate to incised nearly 3/4 of width, distal pinnae often strongly adnate;

segments of rather harsh texture, somewhat oblique, rounded-deltate to often acute;

proximal pair of veins from adjacent segments united below sinus with excurrent vein.

Sori

round, supramedial;

indusia tan, often ciliate;

sporangia glabrous.

round, medial to supramedial;

indusia tan, densely stellate-hairy;

sporangia glabrous.

Indument

abaxially of moderately to densely set hairs to 1 mm on rachises, costae, and veins, glands lacking or yellowish to light orangish, mostly sessile on blade tissue;

blades adaxially often with hairs on veins.

on both surfaces of numerous, sessile or short-stalked, stellate hairs 0.1–0.2 mm on costae, veins, and blade tissue;

rachises and costae sometimes with longer simple hairs to 0.8 mm abaxially.

2n

= 54.

= 144.

Thelypteris noveboracensis

Thelypteris sclerophylla

Habitat Terrestrial in moist woods, especially near swamps, streams, and in vernal seeps of ravines, often in slightly disturbed secondary forests, frequently forming large colonies Terrestrial or on rock in limestone hammocks
Elevation 0–1100 m (0–3600 ft) 0–50 m (0–200 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IL; IN; KY; LA; MA; MD; ME; MI; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WV; NB; NF; NS; ON; PE; QC; SPM
[WildflowerSearch map]
from FNA
FL; West Indies in the Greater Antilles
[BONAP county map]
Discussion

In the flora Thelypteris sclerophylla is known only from Dade County, Florida, where it is rare.

C. Christensen (1913), C. V. Morton (1951), and D. B. Lellinger (1985) have attributed the basionym to Kunze in Sprengel, but Sprengel clearly credited Poeppig, rightly or wrongly. Sprengel's original description also differs in a number of details from that by G. Kunze (Linnaea 9: 92. 1834), so that Kunze's later attribution of the basionym to himself cannot be accepted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Thelypteridaceae > Thelypteris > subg. Parathelypteris Thelypteridaceae > Thelypteris > subg. Goniopteris
Sibling taxa
T. augescens, T. dentata, T. grandis, T. hispidula, T. interrupta, T. kunthii, T. nevadensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. sclerophylla, T. serrata, T. simulata, T. tetragona
T. augescens, T. dentata, T. grandis, T. hispidula, T. interrupta, T. kunthii, T. nevadensis, T. noveboracensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. serrata, T. simulata, T. tetragona
Synonyms Polypodium noveboracense, Dryopteris noveboracensis, Parathelypteris noveboracensis, T. thelypterioides Aspidium sclerophyllum, Dryopteris sclerophylla, Goniopteris sclerophylla
Name authority (Linnaeus) Nieuwland: Amer. Midl. Naturalist 1: 225. (1910) (Poeppig ex Sprengel) C. V. Morton: Amer. Fern J. 41: 87. (1951)
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