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fougère de New York, New York fern, thélyptéride de New York

hairy maiden fern, roughhairy maiden fern

Stems

usually long-creeping, 1.5–2.5 mm diam.

Leaves

monomorphic, dying back in winter, mostly evenly spaced 1 cm or more (sun-gathering leaves in loose cluster), (25–)40–85 cm.

Petiole

straw-colored, 4–25 cm × 1–3 mm, at base with scales tan to reddish brown, ovate, glabrous.

Blade

elliptic, 15–60 cm, proximal 4–10 pinna pairs gradually smaller toward base (smallest often less than 5 mm), blade tapering gradually to pinnatifid apex.

Pinnae

deeply pinnatifid to within 1 mm of costa, 3–9(–13) × 1–2(–2.5) cm;

segments oblong to linear, somewhat oblique, entire to crenulate;

proximal pair of veins from adjacent segments meeting margin above sinus.

Sori

round, supramedial;

indusia tan, often ciliate;

sporangia glabrous.

Indument

abaxially of moderately to densely set hairs to 1 mm on rachises, costae, and veins, glands lacking or yellowish to light orangish, mostly sessile on blade tissue;

blades adaxially often with hairs on veins.

2n

= 54.

Thelypteris noveboracensis

Thelypteris hispidula

Habitat Terrestrial in moist woods, especially near swamps, streams, and in vernal seeps of ravines, often in slightly disturbed secondary forests, frequently forming large colonies
Elevation 0–1100 m (0–3600 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; GA; IL; IN; KY; LA; MA; MD; ME; MI; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WV; NB; NF; NS; ON; PE; QC; SPM
[WildflowerSearch map]
from FNA
AL; FL; GA; LA; MS; SC; TX; North America; Mexico; Central America; South America; tropical and subtropical; West Indies in the Antilles; Asia; Africa
[WildflowerSearch map]
Discussion

Varieties 4 (1 in the flora).

This species and the next are included in Christella subg. Christella by R. E. Holttum (1982).

The relationship between Old World and New World varieties is unstudied.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Thelypteridaceae > Thelypteris > subg. Parathelypteris Thelypteridaceae > Thelypteris > subg. Cyclosorus
Sibling taxa
T. augescens, T. dentata, T. grandis, T. hispidula, T. interrupta, T. kunthii, T. nevadensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. sclerophylla, T. serrata, T. simulata, T. tetragona
T. augescens, T. dentata, T. grandis, T. interrupta, T. kunthii, T. nevadensis, T. noveboracensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. sclerophylla, T. serrata, T. simulata, T. tetragona
Subordinate taxa
T. hispidula var. versicolor
Synonyms Polypodium noveboracense, Dryopteris noveboracensis, Parathelypteris noveboracensis, T. thelypterioides Aspidium hispidulum, Christella hispidula, Cyclosorus quadrangularis, Dryopteris hispidula, Dryopteris quadrangularis, T. quadrangularis
Name authority (Linnaeus) Nieuwland: Amer. Midl. Naturalist 1: 225. (1910) (Decaisne) C. F. Reed: Phytologia 17: 283. (1968)
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