Thelypteris noveboracensis |
Thelypteridaceae |
|||||||||
---|---|---|---|---|---|---|---|---|---|---|
fougère de New York, New York fern, thélyptéride de New York |
marsh fern family, thelypteris family |
|||||||||
Habit | Plants terrestrial or on rock [epiphytic]. | |||||||||
Stems | usually long-creeping, 1.5–2.5 mm diam. |
creeping to erect, scaly at apex. |
||||||||
Leaves | monomorphic, dying back in winter, mostly evenly spaced 1 cm or more (sun-gathering leaves in loose cluster), (25–)40–85 cm. |
monomorphic or somewhat dimorphic [dimorphic]. |
||||||||
Petiole | straw-colored, 4–25 cm × 1–3 mm, at base with scales tan to reddish brown, ovate, glabrous. |
in cross section with 2 crescent-shaped vascular bundles at base. |
||||||||
Blade | elliptic, 15–60 cm, proximal 4–10 pinna pairs gradually smaller toward base (smallest often less than 5 mm), blade tapering gradually to pinnatifid apex. |
pinnate to pinnate-pinnatifid, rarely more than 2-pinnate [simple]; rachis grooved adaxially or not, grooves not continuous with grooves of next order. |
||||||||
Pinnae | deeply pinnatifid to within 1 mm of costa, 3–9(–13) × 1–2(–2.5) cm; segments oblong to linear, somewhat oblique, entire to crenulate; proximal pair of veins from adjacent segments meeting margin above sinus. |
|||||||||
Veins | free or anastomosing, running to margin, areoles with or without included free veinlets. |
|||||||||
Sori | round, supramedial; indusia tan, often ciliate; sporangia glabrous. |
inframedial to supramedial, occasionally nearly marginal, round or oblong, rarely elongate along veins; indusia reniform or sometimes absent. |
||||||||
Spores | bilateral, monolete [rarely globose-tetrahedral and trilete], usually with a prominent, crested, echinate, or reticulate perispore. |
|||||||||
Gametophytes | green, cordate, usually hairy or glandular; antheridia 3-celled. |
|||||||||
Indument | abaxially of moderately to densely set hairs to 1 mm on rachises, costae, and veins, glands lacking or yellowish to light orangish, mostly sessile on blade tissue; blades adaxially often with hairs on veins. |
of transparent, needlelike, hooked, septate, or stellate hairs, or rarely hairs lacking. |
||||||||
2n | = 54. |
|||||||||
Thelypteris noveboracensis |
Thelypteridaceae |
|||||||||
Habitat | Terrestrial in moist woods, especially near swamps, streams, and in vernal seeps of ravines, often in slightly disturbed secondary forests, frequently forming large colonies | |||||||||
Elevation | 0–1100 m (0–3600 ft) | |||||||||
Distribution |
AL; AR; CT; DC; DE; GA; IL; IN; KY; LA; MA; MD; ME; MI; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WV; NB; NF; NS; ON; PE; QC; SPM
|
Mostly tropical |
||||||||
Discussion | Members of Thelypteridaceae have historically been associated with Dryopteridaceae (in particular, Dryopteris) but in fact have no close relationship with that family. Thelypteris and allies differ from Dryopteris and allies by their indument of transparent needlelike hairs (versus needlelike hairs absent in Dryopteridaceae); general absence of blade scales (versus blade scales often present); petiole vasculature in cross section with two crescent-shaped bundles (versus many round bundles arranged in an arc, Athyrium and allies exceptional); generally 1-pinnate to pinnate-pinnatifid blades (versus often more divided); veins usually not forking in the ultimate segments (versus often forking); adaxial grooves discontinuous from rachis to costae, or grooves lacking (versus grooves often continuous); and chromosome base numbers from 27–36 (versus generally 40, 41). Genera 1 to ca. 30, depending on circumscription, species ca. 900 (as circumscribed here, 3 genera and 25 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|||||||||
Key |
|
|||||||||
Source | FNA vol. 2. | FNA vol. 2, p. 206. | ||||||||
Parent taxa | Thelypteridaceae > Thelypteris > subg. Parathelypteris | |||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Polypodium noveboracense, Dryopteris noveboracensis, Parathelypteris noveboracensis, T. thelypterioides | |||||||||
Name authority | (Linnaeus) Nieuwland: Amer. Midl. Naturalist 1: 225. (1910) | Ching ex Pichi Sermolli | ||||||||
Web links |
|