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Nevada Marsh fern, Sierra Marsh fern, Sierra wood fern

Hottentot fern, spready tri-vein fern, Willdenow's fern, Willdenow's maiden fern

Stems

creeping for 2–5 cm, then ascending or suberect, 1.5–3 mm diam.

long-creeping, cordlike, 3–6 mm diam.

Leaves

monomorphic, dying back in winter, tightly clustered, (25–)40–105 cm.

monomorphic, evergreen, 3–6 cm apart, 50–150(–250) cm.

Petiole

straw-colored, 3–20(–35) cm × 1–3 mm, at base with scales tan to reddish brown, ovate, glabrous.

straw-colored to tan, 20–125 cm × 3–6 mm, scaleless.

Blade

elliptic, 20–70 cm, proximal 4–10 pinna pairs gradually reduced (smallest 5–20 mm), blade tapering gradually to pinnatifid apex.

30–125 cm, broadest at base, gradually narrowed distally to pinnatifid apex.

Pinnae

3–10 × (0.6–)1–2 cm, deeply pinnatifid to within 1 mm of costa;

segments oblong to linear, oblique (sides slanted, not perpendicular to costa), entire to crenulate;

proximal pair of veins from adjacent segments meeting margin above sinus.

7–30 × 1–2 cm, incised 1/3–1/2(–3/5) of width;

segments deltate, rounded to acute;

proximal pair of veins from adjacent segments united at acute or obtuse angle below sinus, with excurrent vein 2–4 mm.

Sori

round, supramedial;

indusia tan, glabrous or short-ciliate, sometimes also with glands;

sporangia glabrous.

round, medial to supramedial;

indusia tan, glabrous to hairy;

sporangia with red- or orange-capped, stalked, globose glands arising from sporangial stalks.

Indument

abaxially of sparsely set hairs 0.2–0.7 mm on rachises, costae, and sometimes veins, also of numerous orangish, sessile to usually short-stalked glands on blade tissue;

blades adaxially glabrous except along costae.

abaxially of hairs 0.1–0.3 mm on costae and veins, or hairs often lacking, costae also with tan, ovate scales;

veins, costules, and costae adaxially glabrous or sparsely pubescent;

blade tissue without hairs on both sides, or hairy abaxially, usually with red to orange, shiny, sessile, hemispheric glands abaxially.

2n

= 54.

= 144.

Thelypteris nevadensis

Thelypteris interrupta

Habitat Terrestrial in woods and meadows, especially near springs, seepage areas, and streams Wet roadside ditches, riverbanks, marshes, and cypress swamps
Elevation 0–1800 m (0–5900 ft) 0–50 m (0–200 ft)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
from FNA
FL; LA; Mexico; Central America; West Indies in the Antilles; South America to Argentina; tropical and subtropical Asia; Africa
Discussion

Thelypteris nevadensis is named for the Sierra Nevada and, contrary to its common name, is not found in Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

D. B. Lellinger (1985) applied the name Thelypteris interrupta to specimens from India, while using T. totta (type from South Africa) for North American and South American specimens. Diploid cytotypes are known from Africa and Asia, whereas all counts from the Neotropics are tetraploid. Until more counts are available and the morphologic variation (chiefly in glands, pubescence, and leaf size) in this species complex is better understood, I prefer to circumscribe the species broadly.

R. E. Holttum (1982) circumscribed Cyclosorus (as a genus) to include this species and one or two others.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Thelypteridaceae > Thelypteris > subg. Parathelypteris Thelypteridaceae > Thelypteris > subg. Cyclosorus
Sibling taxa
T. augescens, T. dentata, T. grandis, T. hispidula, T. interrupta, T. kunthii, T. noveboracensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. sclerophylla, T. serrata, T. simulata, T. tetragona
T. augescens, T. dentata, T. grandis, T. hispidula, T. kunthii, T. nevadensis, T. noveboracensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. sclerophylla, T. serrata, T. simulata, T. tetragona
Synonyms Nephrodium nevadense, Dryopteris nevadensis, Dryopteris oregana, Parathelypteris nevadensis Pteris interrupta, Cyclosorus gongylodes, Cyclosorus interruptus, Cyclosorus tottus, Dryopteris gongylodes, T. gongylodes, T. totta
Name authority (Baker) Clute ex C. V. Morton: Amer. Fern J. 48: 139. (1958) (Willdenow) K. Iwatsuki: Jap. J. Bot. 38: 314. (1963)
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