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Nevada Marsh fern, Sierra Marsh fern, Sierra wood fern

abrupt-tip maiden fern

Stems

creeping for 2–5 cm, then ascending or suberect, 1.5–3 mm diam.

creeping, 4–8 mm diam.

Leaves

monomorphic, dying back in winter, tightly clustered, (25–)40–105 cm.

monomorphic, evergreen, (0.5–)1–3.5 cm apart, (30–)65–140 cm.

Petiole

straw-colored, 3–20(–35) cm × 1–3 mm, at base with scales tan to reddish brown, ovate, glabrous.

straw-colored, 15–70 cm × 2–7(–9) mm, at base with scales tan to brownish, linear-lanceolate, hairy at margin.

Blade

elliptic, 20–70 cm, proximal 4–10 pinna pairs gradually reduced (smallest 5–20 mm), blade tapering gradually to pinnatifid apex.

30–70 cm, broadest at or near base, abruptly narrowed distally, apical pinna ± similar to lateral pinnae, 5–17 × 1–3(–5) cm.

Pinnae

3–10 × (0.6–)1–2 cm, deeply pinnatifid to within 1 mm of costa;

segments oblong to linear, oblique (sides slanted, not perpendicular to costa), entire to crenulate;

proximal pair of veins from adjacent segments meeting margin above sinus.

(4–)10–22(–28) × (0.3–)0.7–1.5 cm, incised 1/2–3/4 of width;

segments somewhat curved, margins revolute, those at base of proximal pinnae slightly elongate;

proximal pair of veins from adjacent segments running to sinus.

Sori

round, supramedial;

indusia tan, glabrous or short-ciliate, sometimes also with glands;

sporangia glabrous.

round, medial to supramedial;

indusia tan, bearing hairs 0.2–0.4 mm;

sporangia glabrous.

Indument

abaxially of sparsely set hairs 0.2–0.7 mm on rachises, costae, and sometimes veins, also of numerous orangish, sessile to usually short-stalked glands on blade tissue;

blades adaxially glabrous except along costae.

abaxially of hairs 0.2–0.4 mm on costae, veins, and blade tissue, also of brownish scales 0.6–1.2 mm on costae;

veins and blade tissue glabrous adaxially.

2n

= 54.

= 144.

Thelypteris nevadensis

Thelypteris augescens

Habitat Terrestrial in woods and meadows, especially near springs, seepage areas, and streams Limestone banks, in sun or partial shade
Elevation 0–1800 m (0–5900 ft) 0–50 m (0–200 ft)
Distribution
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
from FNA
FL; s Mexico; West Indies in the Bahamas; Cuba; Central America in Guatemala
Discussion

Thelypteris nevadensis is named for the Sierra Nevada and, contrary to its common name, is not found in Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Thelypteris augescens occasionally hybridizes with T. kunthii and T. ovata var. ovata in southern Florida.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Thelypteridaceae > Thelypteris > subg. Parathelypteris Thelypteridaceae > Thelypteris > subg. Cyclosorus
Sibling taxa
T. augescens, T. dentata, T. grandis, T. hispidula, T. interrupta, T. kunthii, T. noveboracensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. sclerophylla, T. serrata, T. simulata, T. tetragona
T. dentata, T. grandis, T. hispidula, T. interrupta, T. kunthii, T. nevadensis, T. noveboracensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. sclerophylla, T. serrata, T. simulata, T. tetragona
Synonyms Nephrodium nevadense, Dryopteris nevadensis, Dryopteris oregana, Parathelypteris nevadensis Aspidium augescens, Christella augescens, Dryopteris augescens
Name authority (Baker) Clute ex C. V. Morton: Amer. Fern J. 48: 139. (1958) (Link) Munz & I. M. Johnston: Amer. Fern J. 12: 75. (1922)
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