Thelypteris nevadensis |
Thelypteridaceae |
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Nevada Marsh fern, Sierra Marsh fern, Sierra wood fern |
marsh fern family, thelypteris family |
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Habit | Plants terrestrial or on rock [epiphytic]. | |||||||||
Stems | creeping for 2–5 cm, then ascending or suberect, 1.5–3 mm diam. |
creeping to erect, scaly at apex. |
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Leaves | monomorphic, dying back in winter, tightly clustered, (25–)40–105 cm. |
monomorphic or somewhat dimorphic [dimorphic]. |
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Petiole | straw-colored, 3–20(–35) cm × 1–3 mm, at base with scales tan to reddish brown, ovate, glabrous. |
in cross section with 2 crescent-shaped vascular bundles at base. |
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Blade | elliptic, 20–70 cm, proximal 4–10 pinna pairs gradually reduced (smallest 5–20 mm), blade tapering gradually to pinnatifid apex. |
pinnate to pinnate-pinnatifid, rarely more than 2-pinnate [simple]; rachis grooved adaxially or not, grooves not continuous with grooves of next order. |
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Pinnae | 3–10 × (0.6–)1–2 cm, deeply pinnatifid to within 1 mm of costa; segments oblong to linear, oblique (sides slanted, not perpendicular to costa), entire to crenulate; proximal pair of veins from adjacent segments meeting margin above sinus. |
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Veins | free or anastomosing, running to margin, areoles with or without included free veinlets. |
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Sori | round, supramedial; indusia tan, glabrous or short-ciliate, sometimes also with glands; sporangia glabrous. |
inframedial to supramedial, occasionally nearly marginal, round or oblong, rarely elongate along veins; indusia reniform or sometimes absent. |
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Spores | bilateral, monolete [rarely globose-tetrahedral and trilete], usually with a prominent, crested, echinate, or reticulate perispore. |
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Gametophytes | green, cordate, usually hairy or glandular; antheridia 3-celled. |
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Indument | abaxially of sparsely set hairs 0.2–0.7 mm on rachises, costae, and sometimes veins, also of numerous orangish, sessile to usually short-stalked glands on blade tissue; blades adaxially glabrous except along costae. |
of transparent, needlelike, hooked, septate, or stellate hairs, or rarely hairs lacking. |
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2n | = 54. |
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Thelypteris nevadensis |
Thelypteridaceae |
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Habitat | Terrestrial in woods and meadows, especially near springs, seepage areas, and streams | |||||||||
Elevation | 0–1800 m (0–5900 ft) | |||||||||
Distribution |
CA; OR; WA; BC
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Mostly tropical |
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Discussion | Thelypteris nevadensis is named for the Sierra Nevada and, contrary to its common name, is not found in Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Members of Thelypteridaceae have historically been associated with Dryopteridaceae (in particular, Dryopteris) but in fact have no close relationship with that family. Thelypteris and allies differ from Dryopteris and allies by their indument of transparent needlelike hairs (versus needlelike hairs absent in Dryopteridaceae); general absence of blade scales (versus blade scales often present); petiole vasculature in cross section with two crescent-shaped bundles (versus many round bundles arranged in an arc, Athyrium and allies exceptional); generally 1-pinnate to pinnate-pinnatifid blades (versus often more divided); veins usually not forking in the ultimate segments (versus often forking); adaxial grooves discontinuous from rachis to costae, or grooves lacking (versus grooves often continuous); and chromosome base numbers from 27–36 (versus generally 40, 41). Genera 1 to ca. 30, depending on circumscription, species ca. 900 (as circumscribed here, 3 genera and 25 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 2. | FNA vol. 2, p. 206. | ||||||||
Parent taxa | Thelypteridaceae > Thelypteris > subg. Parathelypteris | |||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Nephrodium nevadense, Dryopteris nevadensis, Dryopteris oregana, Parathelypteris nevadensis | |||||||||
Name authority | (Baker) Clute ex C. V. Morton: Amer. Fern J. 48: 139. (1958) | Ching ex Pichi Sermolli | ||||||||
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