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Hottentot fern, spready tri-vein fern, Willdenow's fern, Willdenow's maiden fern

Nevada Marsh fern, Sierra Marsh fern, Sierra wood fern

Stems

long-creeping, cordlike, 3–6 mm diam.

creeping for 2–5 cm, then ascending or suberect, 1.5–3 mm diam.

Leaves

monomorphic, evergreen, 3–6 cm apart, 50–150(–250) cm.

monomorphic, dying back in winter, tightly clustered, (25–)40–105 cm.

Petiole

straw-colored to tan, 20–125 cm × 3–6 mm, scaleless.

straw-colored, 3–20(–35) cm × 1–3 mm, at base with scales tan to reddish brown, ovate, glabrous.

Blade

30–125 cm, broadest at base, gradually narrowed distally to pinnatifid apex.

elliptic, 20–70 cm, proximal 4–10 pinna pairs gradually reduced (smallest 5–20 mm), blade tapering gradually to pinnatifid apex.

Pinnae

7–30 × 1–2 cm, incised 1/3–1/2(–3/5) of width;

segments deltate, rounded to acute;

proximal pair of veins from adjacent segments united at acute or obtuse angle below sinus, with excurrent vein 2–4 mm.

3–10 × (0.6–)1–2 cm, deeply pinnatifid to within 1 mm of costa;

segments oblong to linear, oblique (sides slanted, not perpendicular to costa), entire to crenulate;

proximal pair of veins from adjacent segments meeting margin above sinus.

Sori

round, medial to supramedial;

indusia tan, glabrous to hairy;

sporangia with red- or orange-capped, stalked, globose glands arising from sporangial stalks.

round, supramedial;

indusia tan, glabrous or short-ciliate, sometimes also with glands;

sporangia glabrous.

Indument

abaxially of hairs 0.1–0.3 mm on costae and veins, or hairs often lacking, costae also with tan, ovate scales;

veins, costules, and costae adaxially glabrous or sparsely pubescent;

blade tissue without hairs on both sides, or hairy abaxially, usually with red to orange, shiny, sessile, hemispheric glands abaxially.

abaxially of sparsely set hairs 0.2–0.7 mm on rachises, costae, and sometimes veins, also of numerous orangish, sessile to usually short-stalked glands on blade tissue;

blades adaxially glabrous except along costae.

2n

= 144.

= 54.

Thelypteris interrupta

Thelypteris nevadensis

Habitat Wet roadside ditches, riverbanks, marshes, and cypress swamps Terrestrial in woods and meadows, especially near springs, seepage areas, and streams
Elevation 0–50 m (0–200 ft) 0–1800 m (0–5900 ft)
Distribution
from FNA
FL; LA; Mexico; Central America; West Indies in the Antilles; South America to Argentina; tropical and subtropical Asia; Africa
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
Discussion

D. B. Lellinger (1985) applied the name Thelypteris interrupta to specimens from India, while using T. totta (type from South Africa) for North American and South American specimens. Diploid cytotypes are known from Africa and Asia, whereas all counts from the Neotropics are tetraploid. Until more counts are available and the morphologic variation (chiefly in glands, pubescence, and leaf size) in this species complex is better understood, I prefer to circumscribe the species broadly.

R. E. Holttum (1982) circumscribed Cyclosorus (as a genus) to include this species and one or two others.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Thelypteris nevadensis is named for the Sierra Nevada and, contrary to its common name, is not found in Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Thelypteridaceae > Thelypteris > subg. Cyclosorus Thelypteridaceae > Thelypteris > subg. Parathelypteris
Sibling taxa
T. augescens, T. dentata, T. grandis, T. hispidula, T. kunthii, T. nevadensis, T. noveboracensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. sclerophylla, T. serrata, T. simulata, T. tetragona
T. augescens, T. dentata, T. grandis, T. hispidula, T. interrupta, T. kunthii, T. noveboracensis, T. ovata, T. palustris, T. patens, T. pilosa, T. puberula, T. quelpaertensis, T. reptans, T. resinifera, T. reticulata, T. sclerophylla, T. serrata, T. simulata, T. tetragona
Synonyms Pteris interrupta, Cyclosorus gongylodes, Cyclosorus interruptus, Cyclosorus tottus, Dryopteris gongylodes, T. gongylodes, T. totta Nephrodium nevadense, Dryopteris nevadensis, Dryopteris oregana, Parathelypteris nevadensis
Name authority (Willdenow) K. Iwatsuki: Jap. J. Bot. 38: 314. (1963) (Baker) Clute ex C. V. Morton: Amer. Fern J. 48: 139. (1958)
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