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Wright's stanleyella, Wright's thelypody

Rollins' thelypody

Habit Biennials; slightly glaucous, glabrous. Biennials; glaucous, glabrous throughout.
Stems

branched distally, (6.5–)9.5–22.5(–28) dm.

branched distally, (4–)6–16(–20) dm.

Basal leaves

(and proximal cauline): petiole (2–)4.5–8.5(–13.7) cm;

blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5–)9.5–22.5(–28) cm × 28–55(–75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate).

petiole 0.5–1.3(–1.5) cm, not ciliate;

blade spatulate or oblanceolate to obovate, 1.3–4(–7) cm × (4–)6–13(–18) mm, margins usually dentate to repand, rarely entire.

Cauline leaves

petiolate;

blade lanceolate to linear-lanceolate, 3.5–8.5(–13.5) cm × 7–14(–23) mm, margins usually entire or dentate, rarely lobed.

(erect, appressed to stem at least partly);

sessile;

blade linear to linear-lanceolate, (1.8–)1.3–4.5(–6) cm × (1–)1.7–7(–8) mm, (base auriculate or sagittate), margins entire.

Racemes

somewhat lax, considerably elongated in fruit.

dense, terminal not elongated in fruit, (flower buds narrowly oblong).

Flowers

sepals spreading or ascending oblong to linear, (3–)4–6(–7) × (0.8–)1–1.3(–1.5) mm;

petals usually white, rarely lavender, linear or oblong, 4–7.5(–9) × 1–1.8(–2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base);

nectar glands confluent;

filaments equal, (3–)3.5–6.5(–8.5) mm;

anthers (1–)1.5–2.5(–3) mm, circinately coiled;

gynophore 0.2–2(–5) mm.

sepals erect, usually linear to linear-oblong, rarely oblong, 4–6(–7) × (0.8–)1–1.5 mm;

petals lavender to purple, spatulate to obovate or oblong, (6–)7–9.5(–12) × (1.2–)1.5–2(–3) mm, margins not crisped, claw differentiated from blade, [slender, 3–4.5(–5.5) mm, narrowest at base];

nectar glands confluent, subtending bases of stamens;

filaments tetradynamous, median pairs (5–)5.5–7.5(–8.5) mm, lateral pair (4–)5–6.5(7.5) mm;

anthers exserted, linear to linear-oblong, 2–3(–4) mm, not circinately coiled;

gynophore 0.5–2.5(–6) mm.

Fruiting pedicels

horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5–)7–13(–17) mm, slightly flattened at base.

usually horizontal to divaricate, rarely divaricate-ascending, straight, slender, (6–)7–14(–18) mm, often not flattened at base.

Fruits

horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5–)3.8–7.4(–9) cm × 1–1.2(–1.5) mm, (replum not constricted between seeds);

ovules 76–128 per ovary;

style usually clavate to subclavate, rarely cylindrical, (0.5–)0.8–2(–3) mm.

erect distally, slightly torulose, usually strongly incurved, (overtopping buds), rarely straight and divaricate-ascending, terete, (1.8–)2.8–5(–6.3) cm × 0.7–1 mm;

ovules 52–70 per ovary;

style cylindrical, (0.5–)0.8–1.5(–2) mm.

Seeds

0.7–1.3(–1.5) × 0.5–0.8 mm.

1–1.3(–1.5) × 0.5–0.6(–0.8) mm.

2n

= 26.

Thelypodium wrightii

Thelypodium rollinsii

Phenology Flowering (Mar-)Jun–Oct. Flowering Jun–Aug.
Habitat Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands Alkaline flats
Elevation 1200-2300 m (3900-7500 ft) 1500-2000 m (4900-6600 ft)
Distribution
from FNA
AZ; CO; NM; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Hidalgo)
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp. oklahomensis and subsp. wrightii, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Thelypodium rollinsii is restricted mostly to Beaver, Carbon, Juab, Millard, Piute, Sanpete, and Sevier counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 738. FNA vol. 7, p. 736.
Parent taxa Brassicaceae > tribe Thelypodieae > Thelypodium Brassicaceae > tribe Thelypodieae > Thelypodium
Sibling taxa
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
Synonyms Stanleyella wrightii, T. wrightii subsp. oklahomensis
Name authority A. Gray: Smithsonian Contr. Knowl. 3(5): 7. (1852) Al-Shehbaz: Contr. Gray Herb. 204: 97, plates, figs. (1973)
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