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Wright's stanleyella, Wright's thelypody

cut-leaf thelypody, feathery thelypodium, many-flower thelypody, thick-leaf thelypody, thick-leaf thelypody (var. laciniatum), western thelypodium

Habit Biennials; slightly glaucous, glabrous. Biennials; somewhat glaucous, glabrous throughout.
Stems

branched distally, (6.5–)9.5–22.5(–28) dm.

branched proximally and/or distally, (1.3–)2.6–10(–14.5) dm.

Basal leaves

(and proximal cauline): petiole (2–)4.5–8.5(–13.7) cm;

blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5–)9.5–22.5(–28) cm × 28–55(–75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate).

(and proximal cauline): petiole (1–)1.5–10(–15) cm;

blade lanceolate to deltate-lanceolate, or oblong to ovate (lateral lobes oblong to ovate), (4.3–)6.8–24(–45) cm × (10–)18–85(–140) mm, margins pinnately lobed or laciniate (lobes sinuate, laciniate, or dentate).

Cauline leaves

petiolate;

blade lanceolate to linear-lanceolate, 3.5–8.5(–13.5) cm × 7–14(–23) mm, margins usually entire or dentate, rarely lobed.

petiolate;

blade lanceolate to linear-lanceolate, similar to basal, smaller distally, margins entire or sinuate to laciniate.

Racemes

somewhat lax, considerably elongated in fruit.

dense, slightly elongated in fruit (to 9.5 dm, flower buds narrowly oblong).

Flowers

sepals spreading or ascending oblong to linear, (3–)4–6(–7) × (0.8–)1–1.3(–1.5) mm;

petals usually white, rarely lavender, linear or oblong, 4–7.5(–9) × 1–1.8(–2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base);

nectar glands confluent;

filaments equal, (3–)3.5–6.5(–8.5) mm;

anthers (1–)1.5–2.5(–3) mm, circinately coiled;

gynophore 0.2–2(–5) mm.

sepals erect, oblong to linear-oblong, (3.5–)4–7.5(–9.5) × (0.8–)1–2(–2.5) mm;

petals white or purple, often linear, sometimes linear-lanceolate, (6–)8.5–18(–20) × 0.3–0.8(–1.5) mm, margins slightly crisped, claw strongly differentiated from blade, [(2.5–)3–6(–7) mm, widest at base];

nectar glands confluent, lateral and median;

filaments subequal, (4.5–)6.5–12(–15) mm;

anthers exserted, linear, 2–4(–5.5) mm, circinately coiled, (apiculate);

gynophore (0.5–)1–5(–8.5) mm.

Fruiting pedicels

horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5–)7–13(–17) mm, slightly flattened at base.

horizontal to, rarely, divaricate, straight, often stout, (2–)3–6.5(–15) mm, often flattened at base.

Fruits

horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5–)3.8–7.4(–9) cm × 1–1.2(–1.5) mm, (replum not constricted between seeds);

ovules 76–128 per ovary;

style usually clavate to subclavate, rarely cylindrical, (0.5–)0.8–2(–3) mm.

divaricate-ascending to slightly reflexed, torulose, somewhat tortuous, terete, (2.5–)3.5–10(–12) cm × (0.7–)1–1.5(–2) mm;

ovules 56–108 per ovary;

style usually cylindrical, rarely subclavate, (0.3–)0.7–2.5(–4) mm.

Seeds

0.7–1.3(–1.5) × 0.5–0.8 mm.

(0.7–)1–1.5(–1.8) × 0.5–0.7(–1) mm.

2n

= 26.

Thelypodium wrightii

Thelypodium laciniatum

Phenology Flowering (Mar-)Jun–Oct. Flowering Apr–Aug.
Habitat Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands Rock crevices, cliffs, rocky outcrops, among boulders, serpentine rock, talus, canyon walls, limestone ledges
Elevation 1200-2300 m (3900-7500 ft) 0-2400 m (0-7900 ft)
Distribution
from FNA
AZ; CO; NM; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Hidalgo)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; NV; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp. oklahomensis and subsp. wrightii, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 738. FNA vol. 7, p. 734.
Parent taxa Brassicaceae > tribe Thelypodieae > Thelypodium Brassicaceae > tribe Thelypodieae > Thelypodium
Sibling taxa
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
Synonyms Stanleyella wrightii, T. wrightii subsp. oklahomensis Macropodium laciniatum, Pachypodium laciniatum, T. laciniatum var. streptanthoides, T. leptosepalum, T. streptanthoides
Name authority A. Gray: Smithsonian Contr. Knowl. 3(5): 7. (1852) (Hooker) Endlicher: in W. G. Walpers, Repert. Bot. Syst. 1: 172. (1842)
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