Thelypodium wrightii |
Thelypodium integrifolium |
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Wright's stanleyella, Wright's thelypody |
entire leaf thelypodium, entire-leaf thelypody |
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Habit | Biennials; slightly glaucous, glabrous. | Biennials; somewhat glaucous throughout, glabrous. | ||||||||||||||||
Stems | branched distally, (6.5–)9.5–22.5(–28) dm. |
paniculately branched distally, (2–)4.5–17(–28) dm. |
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Basal leaves | (and proximal cauline): petiole (2–)4.5–8.5(–13.7) cm; blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5–)9.5–22.5(–28) cm × 28–55(–75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate). |
petiole (0.5–)1.2–10(–15) cm; blade usually oblong to lanceolate, or oblanceolate to spatulate, rarely ovate or obovate, (3.7–)5–31(–54) cm × (12–)16–78(–140) mm, margins usually entire, rarely dentate or repand. |
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Cauline leaves | petiolate; blade lanceolate to linear-lanceolate, 3.5–8.5(–13.5) cm × 7–14(–23) mm, margins usually entire or dentate, rarely lobed. |
(ascending, not appressed to stem); sessile; blade linear to linear-lanceolate, (1–)2.2–8.3(–19.5) cm × (2–)3–11(–25) mm, (base cuneate to attenuate, not auriculate), margins entire. |
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Racemes | somewhat lax, considerably elongated in fruit. |
dense, slightly or considerably elongated in fruit, (flower buds narrowly oblong). |
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Flowers | sepals spreading or ascending oblong to linear, (3–)4–6(–7) × (0.8–)1–1.3(–1.5) mm; petals usually white, rarely lavender, linear or oblong, 4–7.5(–9) × 1–1.8(–2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base); nectar glands confluent; filaments equal, (3–)3.5–6.5(–8.5) mm; anthers (1–)1.5–2.5(–3) mm, circinately coiled; gynophore 0.2–2(–5) mm. |
sepals erect, linear to linear-oblong, (3–)3.5–5.5(–7.5) × (0.5–)0.8–1(–1.5) mm; petals white or lavender to purple, spatulate to oblanceolate, (4.5–)6–9(–13) × (0.5–)0.8–1.5(–2.3) mm, margins not crisped, claw differentiated from blade, [slender, (2–)3–4.5(–7) mm, narrowest at base]; nectar glands lateral, flat or toothlike, median glands absent; filaments subequal to slightly tetradynamous, (3–)4.5–7.5(–12.5) mm; anthers exserted, linear to narrowly oblong, (1–)1.5–2.5(–4.5) mm, not circinately coiled; gynophore 0.5–2(–5.5) mm. |
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Fruiting pedicels | horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5–)7–13(–17) mm, slightly flattened at base. |
usually horizontal to divaricate, rarely ascending or reflexed, usually straight, rarely incurved, slender or stout, (2–)3–9(–13) mm, often strongly flattened, rarely not flattened at base. |
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Fruits | horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5–)3.8–7.4(–9) cm × 1–1.2(–1.5) mm, (replum not constricted between seeds); ovules 76–128 per ovary; style usually clavate to subclavate, rarely cylindrical, (0.5–)0.8–2(–3) mm. |
usually horizontal to divaricate-ascending, or ascending, rarely reflexed, usually torulose, rarely submoniliform, usually strongly incurved, sometimes straight, rarely arcuate, terete, (0.8–)1.4–6.5(–8) cm × 1–1.3(–2) mm; ovules 14–40 per ovary; style usually cylindrical, rarely subclavate, 0.5–1.3(–3.2) mm. |
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Seeds | 0.7–1.3(–1.5) × 0.5–0.8 mm. |
(1–)1.2–2(–2.3) × 0.7–1(–1.3) mm. |
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Thelypodium wrightii |
Thelypodium integrifolium |
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Phenology | Flowering (Mar-)Jun–Oct. | |||||||||||||||||
Habitat | Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands | |||||||||||||||||
Elevation | 1200-2300 m (3900-7500 ft) | |||||||||||||||||
Distribution |
AZ; CO; NM; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Hidalgo)
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AZ; CA; CO; ID; MT; ND; NE; NM; NV; OR; SD; UT; WA; WY
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Discussion | Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp. oklahomensis and subsp. wrightii, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 5 (5 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 738. | FNA vol. 7, p. 732. | ||||||||||||||||
Parent taxa | Brassicaceae > tribe Thelypodieae > Thelypodium | Brassicaceae > tribe Thelypodieae > Thelypodium | ||||||||||||||||
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Subordinate taxa | ||||||||||||||||||
Synonyms | Stanleyella wrightii, T. wrightii subsp. oklahomensis | Pachypodium integrifolium, Pleurophragma integrifolium | ||||||||||||||||
Name authority | A. Gray: Smithsonian Contr. Knowl. 3(5): 7. (1852) | (Nuttall) Endlicher: in W. G. Walpers, Repert. Bot. Syst. 1: 172. (1842) | ||||||||||||||||
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