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Wright's stanleyella, Wright's thelypody

entire leaf thelypodium, entire-leaf thelypody

Habit Biennials; slightly glaucous, glabrous. Biennials; somewhat glaucous throughout, glabrous.
Stems

branched distally, (6.5–)9.5–22.5(–28) dm.

paniculately branched distally, (2–)4.5–17(–28) dm.

Basal leaves

(and proximal cauline): petiole (2–)4.5–8.5(–13.7) cm;

blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5–)9.5–22.5(–28) cm × 28–55(–75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate).

petiole (0.5–)1.2–10(–15) cm;

blade usually oblong to lanceolate, or oblanceolate to spatulate, rarely ovate or obovate, (3.7–)5–31(–54) cm × (12–)16–78(–140) mm, margins usually entire, rarely dentate or repand.

Cauline leaves

petiolate;

blade lanceolate to linear-lanceolate, 3.5–8.5(–13.5) cm × 7–14(–23) mm, margins usually entire or dentate, rarely lobed.

(ascending, not appressed to stem);

sessile;

blade linear to linear-lanceolate, (1–)2.2–8.3(–19.5) cm × (2–)3–11(–25) mm, (base cuneate to attenuate, not auriculate), margins entire.

Racemes

somewhat lax, considerably elongated in fruit.

dense, slightly or considerably elongated in fruit, (flower buds narrowly oblong).

Flowers

sepals spreading or ascending oblong to linear, (3–)4–6(–7) × (0.8–)1–1.3(–1.5) mm;

petals usually white, rarely lavender, linear or oblong, 4–7.5(–9) × 1–1.8(–2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base);

nectar glands confluent;

filaments equal, (3–)3.5–6.5(–8.5) mm;

anthers (1–)1.5–2.5(–3) mm, circinately coiled;

gynophore 0.2–2(–5) mm.

sepals erect, linear to linear-oblong, (3–)3.5–5.5(–7.5) × (0.5–)0.8–1(–1.5) mm;

petals white or lavender to purple, spatulate to oblanceolate, (4.5–)6–9(–13) × (0.5–)0.8–1.5(–2.3) mm, margins not crisped, claw differentiated from blade, [slender, (2–)3–4.5(–7) mm, narrowest at base];

nectar glands lateral, flat or toothlike, median glands absent;

filaments subequal to slightly tetradynamous, (3–)4.5–7.5(–12.5) mm;

anthers exserted, linear to narrowly oblong, (1–)1.5–2.5(–4.5) mm, not circinately coiled;

gynophore 0.5–2(–5.5) mm.

Fruiting pedicels

horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5–)7–13(–17) mm, slightly flattened at base.

usually horizontal to divaricate, rarely ascending or reflexed, usually straight, rarely incurved, slender or stout, (2–)3–9(–13) mm, often strongly flattened, rarely not flattened at base.

Fruits

horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5–)3.8–7.4(–9) cm × 1–1.2(–1.5) mm, (replum not constricted between seeds);

ovules 76–128 per ovary;

style usually clavate to subclavate, rarely cylindrical, (0.5–)0.8–2(–3) mm.

usually horizontal to divaricate-ascending, or ascending, rarely reflexed, usually torulose, rarely submoniliform, usually strongly incurved, sometimes straight, rarely arcuate, terete, (0.8–)1.4–6.5(–8) cm × 1–1.3(–2) mm;

ovules 14–40 per ovary;

style usually cylindrical, rarely subclavate, 0.5–1.3(–3.2) mm.

Seeds

0.7–1.3(–1.5) × 0.5–0.8 mm.

(1–)1.2–2(–2.3) × 0.7–1(–1.3) mm.

Thelypodium wrightii

Thelypodium integrifolium

Phenology Flowering (Mar-)Jun–Oct.
Habitat Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands
Elevation 1200-2300 m (3900-7500 ft)
Distribution
from FNA
AZ; CO; NM; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Hidalgo)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; ND; NE; NM; NV; OR; SD; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp. oklahomensis and subsp. wrightii, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 5 (5 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals white; fruiting pedicels whitish, stout, (5-) 6-11(-13) mm
→ 2
1. Petals usually lavender to purple, rarely white; fruiting pedicels not whitish, slender and 2-10 (-13) mm, or stout and 2-5(-6) mm
→ 3
2. Petals (5-)6-9(-10) mm; fruiting pedicels horizontal to divaricate, (5-)6-9(-13) mm; fruits (1.4-)1.9-3.7(-4.5) cm; gynophores (0.5-)1-3 mm; se California, s Nevada, sw Utah.
subsp. affine
2. Petals (7.5-)8-11(-13) mm; fruiting pedicels usually horizontal to reflexed, rarely divaricate, (6-)7-11(-13) mm; fruits (2.2-) 3.5-6.5(-8) cm; gynophores (1-)1.5-4 (-5.5) mm; Grand Canyon, Colorado River, Arizona.
subsp. longicarpum
3. Fruiting pedicels often slender, sometimes stout, not or slightly flattened at base, (4-)6- 10 (-13) mm.
subsp. integrifolium
3. Fruiting pedicels stout, strongly flattened at base, 2-5(-6) mm
→ 4
4. Racemes often strongly congested, central rachises (0.6-)1.2-7(-9) cm; gynophores stout, 0.5-1(-3) mm; e California, Nevada, s Oregon, nw Utah.
subsp. complanatum
4. Racemes somewhat congested, central rachises (3-)8-20(-28) cm; gynophores slender, (0.5-)0.8-3(-4) mm; ne Arizona, w Colorado, nw New Mexico, e Utah.
subsp. gracilipes
Source FNA vol. 7, p. 738. FNA vol. 7, p. 732.
Parent taxa Brassicaceae > tribe Thelypodieae > Thelypodium Brassicaceae > tribe Thelypodieae > Thelypodium
Sibling taxa
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
Subordinate taxa
T. integrifolium subsp. affine, T. integrifolium subsp. complanatum, T. integrifolium subsp. gracilipes, T. integrifolium subsp. integrifolium, T. integrifolium subsp. longicarpum
Synonyms Stanleyella wrightii, T. wrightii subsp. oklahomensis Pachypodium integrifolium, Pleurophragma integrifolium
Name authority A. Gray: Smithsonian Contr. Knowl. 3(5): 7. (1852) (Nuttall) Endlicher: in W. G. Walpers, Repert. Bot. Syst. 1: 172. (1842)
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