Thelypodium wrightii |
Thelypodium flexuosum |
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Wright's stanleyella, Wright's thelypody |
nodding thelypodium, nodding thelypody, spreading thelypodium, spreading thelypody, zigzag thelypody |
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Habit | Biennials; slightly glaucous, glabrous. | Perennials; (caudex woody, covered with persistent petiolar remains); somewhat glaucous, glabrous throughout. |
Stems | branched distally, (6.5–)9.5–22.5(–28) dm. |
(often subdecumbent), branched basally, (flexuous), 1.5–5.6(–8.5) dm. |
Basal leaves | (and proximal cauline): petiole (2–)4.5–8.5(–13.7) cm; blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5–)9.5–22.5(–28) cm × 28–55(–75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate). |
petiole 1–6.5(–13) cm; blade often lanceolate, sometimes oblong or oblanceolate, (2–)3.5–16.5(–20.5) cm × (5–)10–25(–45) mm, margins entire. |
Cauline leaves | petiolate; blade lanceolate to linear-lanceolate, 3.5–8.5(–13.5) cm × 7–14(–23) mm, margins usually entire or dentate, rarely lobed. |
(ascending); sessile; blade lanceolate to linear, 1–7(–11) cm × 2–7(–14) mm, (base sagittate to somewhat amplexicaul), margins entire. |
Racemes | somewhat lax, considerably elongated in fruit. |
(few-flowered, corymbose), elongated in fruit, (flower buds oblong). |
Flowers | sepals spreading or ascending oblong to linear, (3–)4–6(–7) × (0.8–)1–1.3(–1.5) mm; petals usually white, rarely lavender, linear or oblong, 4–7.5(–9) × 1–1.8(–2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base); nectar glands confluent; filaments equal, (3–)3.5–6.5(–8.5) mm; anthers (1–)1.5–2.5(–3) mm, circinately coiled; gynophore 0.2–2(–5) mm. |
sepals erect, oblong, 3–4(–4.5) × 1–1.5(–1.7) mm; petals lavender or white, often spatulate, sometimes oblanceolate or obovate, 6–9(–10) × (1.5–)2–3(–3.5) mm, margins not crisped, claw strongly differentiated from blade, (slender, 2–3.5(–4) mm, narrowest at base); nectar glands lateral, median glands absent; filaments tetradynamous, median pairs 3–4(–5) mm, lateral pair 2–3.5(–4) mm; anthers included, oblong, 1–2(–2.5) mm, not circinately coiled; gynophore (stout), 0.5–1 mm. |
Fruiting pedicels | horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5–)7–13(–17) mm, slightly flattened at base. |
usually horizontal to divaricate, rarely divaricate-ascending, straight or slightly curved upward, slender, (2.5–)4–9(–16) mm, slightly flattened at base. |
Fruits | horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5–)3.8–7.4(–9) cm × 1–1.2(–1.5) mm, (replum not constricted between seeds); ovules 76–128 per ovary; style usually clavate to subclavate, rarely cylindrical, (0.5–)0.8–2(–3) mm. |
erect to ascending, torulose, slightly incurved or straight, terete, 1–2.5(–4.2) cm × 0.8–1(–1.5) mm; ovules 12–30 per ovary; style cylindrical, (0.3–)1–2(–3) mm. |
Seeds | 0.7–1.3(–1.5) × 0.5–0.8 mm. |
(1–)1.3–1.5 × 0.5–8(–1) mm. |
2n | = 26. |
|
Thelypodium wrightii |
Thelypodium flexuosum |
|
Phenology | Flowering (Mar-)Jun–Oct. | Flowering Apr–Jun. |
Habitat | Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands | Often tangled among woody shrubs in moderately to strongly alkaline sandy loam or clay, open deserts |
Elevation | 1200-2300 m (3900-7500 ft) | 1000-2400 m (3300-7900 ft) |
Distribution |
AZ; CO; NM; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Hidalgo)
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CA; ID; NV; OR
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Discussion | Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp. oklahomensis and subsp. wrightii, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 738. | FNA vol. 7, p. 731. |
Parent taxa | Brassicaceae > tribe Thelypodieae > Thelypodium | Brassicaceae > tribe Thelypodieae > Thelypodium |
Sibling taxa | ||
Synonyms | Stanleyella wrightii, T. wrightii subsp. oklahomensis | |
Name authority | A. Gray: Smithsonian Contr. Knowl. 3(5): 7. (1852) | B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1(1,1): 175. (1895) |
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