Thelypodium wrightii |
Thelypodium eucosmum |
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Wright's stanleyella, Wright's thelypody |
arrow-leaf thelypody, world thelypody |
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Habit | Biennials; slightly glaucous, glabrous. | Biennials or perennials; (short-lived); glaucous, glabrous (except petioles). |
Stems | branched distally, (6.5–)9.5–22.5(–28) dm. |
branched distally, 2–10 dm. |
Basal leaves | (and proximal cauline): petiole (2–)4.5–8.5(–13.7) cm; blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5–)9.5–22.5(–28) cm × 28–55(–75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate). |
petiole (0.9–)1.4–3(–4.5) cm, ciliate; blade usually oblanceolate to oblong or lanceolate, rarely ovate or elliptic, (2.8–)3.5–8.8(–11) cm × (7–)10–25(–35) mm, margins often entire or repand, sometimes sinuate. |
Cauline leaves | petiolate; blade lanceolate to linear-lanceolate, 3.5–8.5(–13.5) cm × 7–14(–23) mm, margins usually entire or dentate, rarely lobed. |
(ascending); sessile; blade lanceolate to oblong, 1.8–4.5(–6) cm × 5–16(–24) mm, (base amplexicaul to strongly auriculate), margins entire. |
Racemes | somewhat lax, considerably elongated in fruit. |
lax, slightly elongated in fruit, (flower buds oblong-linear). |
Flowers | sepals spreading or ascending oblong to linear, (3–)4–6(–7) × (0.8–)1–1.3(–1.5) mm; petals usually white, rarely lavender, linear or oblong, 4–7.5(–9) × 1–1.8(–2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base); nectar glands confluent; filaments equal, (3–)3.5–6.5(–8.5) mm; anthers (1–)1.5–2.5(–3) mm, circinately coiled; gynophore 0.2–2(–5) mm. |
sepals erect, linear-oblong, 5–7(–8) × (0.8–)1–1.5(–1.8) mm; petals dark purple, spatulate to oblanceolate, (6.6–)7.5–10(–11.5) × 1–1.8(–2) mm, margins not crisped, claw differentiated from blade, [slender, (3–)3.5–5(–5.5) mm, narrowest at base]; nectar glands confluent, subtending bases of stamens; filaments subequal, (4.5–)6–9(–10) mm; anthers exserted, linear to narrowly oblong, 2.5–4(–4.5) mm, circinately coiled; gynophore (1–)2.5–6(–7.5) mm. |
Fruiting pedicels | horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5–)7–13(–17) mm, slightly flattened at base. |
horizontal to divaricate, usually straight, rarely slightly incurved, slender, (2.5–)3–5.5(–6.5) mm, slightly flattened at base. |
Fruits | horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5–)3.8–7.4(–9) cm × 1–1.2(–1.5) mm, (replum not constricted between seeds); ovules 76–128 per ovary; style usually clavate to subclavate, rarely cylindrical, (0.5–)0.8–2(–3) mm. |
divaricate to ascending, torulose, straight or slightly incurved, terete, (2–)2.4–5(–6.5) cm × 0.7–1(–1.3) mm; ovules 44–58 per ovary; style usually cylindrical, rarely subclavate, 0.5–1.5(–2) mm. |
Seeds | 0.7–1.3(–1.5) × 0.5–0.8 mm. |
0.7–1.5 × 0.5–0.8 mm. |
Thelypodium wrightii |
Thelypodium eucosmum |
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Phenology | Flowering (Mar-)Jun–Oct. | Flowering May–Jul. |
Habitat | Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands | Shady slopes and canyons, pinyon-juniper and oak woodland communities, stream beds, streamsides |
Elevation | 1200-2300 m (3900-7500 ft) | 700-1000 m (2300-3300 ft) |
Distribution |
AZ; CO; NM; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Hidalgo)
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OR
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Discussion | Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp. oklahomensis and subsp. wrightii, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Thelypodium eucosmum is known only from Baker, Grant, and Wheeler counties. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 738. | FNA vol. 7, p. 731. |
Parent taxa | Brassicaceae > tribe Thelypodieae > Thelypodium | Brassicaceae > tribe Thelypodieae > Thelypodium |
Sibling taxa | ||
Synonyms | Stanleyella wrightii, T. wrightii subsp. oklahomensis | |
Name authority | A. Gray: Smithsonian Contr. Knowl. 3(5): 7. (1852) | B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1(1,1): 175. (1895) |
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