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Wright's stanleyella, Wright's thelypody

crisp thelypody, crisped thelypody, wavy leaf thelypodium

Habit Biennials; slightly glaucous, glabrous. Biennials or perennials; (short-lived); often glaucous, glabrous or pubescent.
Stems

branched distally, (6.5–)9.5–22.5(–28) dm.

branched, (0.9–)1.3–7(–12) dm, (somewhat glaucous, glabrous throughout or pubescent basally).

Basal leaves

(and proximal cauline): petiole (2–)4.5–8.5(–13.7) cm;

blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5–)9.5–22.5(–28) cm × 28–55(–75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate).

petiole (0.7–)2–8.5 cm, ciliate;

blade usually oblanceolate to spatulate, rarely obovate or lanceolate, (1.5–)2.2–15(–25) cm × (6–)10–35(–50) mm, margins often pinnately lobed to lyrate, sometimes dentate or entire.

Cauline leaves

petiolate;

blade lanceolate to linear-lanceolate, 3.5–8.5(–13.5) cm × 7–14(–23) mm, margins usually entire or dentate, rarely lobed.

sessile;

blade linear to lanceolate, smaller distally, (base sagittate or auriculate, appressed to stem), margins usually entire, rarely dentate.

Racemes

somewhat lax, considerably elongated in fruit.

dense, elongated in fruit, (flower buds narrowly oblong).

Flowers

sepals spreading or ascending oblong to linear, (3–)4–6(–7) × (0.8–)1–1.3(–1.5) mm;

petals usually white, rarely lavender, linear or oblong, 4–7.5(–9) × 1–1.8(–2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base);

nectar glands confluent;

filaments equal, (3–)3.5–6.5(–8.5) mm;

anthers (1–)1.5–2.5(–3) mm, circinately coiled;

gynophore 0.2–2(–5) mm.

sepals erect to ascending, oblong to linear-oblong, (3–)3.5–6(–8) × (0.8–)1–1.8(–2) mm;

petals white to lavender, linear to narrowly oblanceolate, (6–)6.5–11(–14.5) × 0.5–0.7(–1) mm, margins strongly crisped, claw differentiated from blade, (slender, 2–4 mm, narrowest at base);

nectar glands surrounding bases of lateral stamens, median glands absent;

filaments subequal, (2.5–)4.5–8.5(–10.5) mm;

anthers exserted, linear to narrowly oblong, 2–3.5(–4.5) mm, circinately coiled, (apiculate);

gynophore 0.5–1.5(–3.5) mm.

Fruiting pedicels

horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5–)7–13(–17) mm, slightly flattened at base.

erect to erect-ascending, (appressed to rachis at least basally), straight or slightly curved, slender, (1.5–)2–5(–10) mm, not flattened at base.

Fruits

horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5–)3.8–7.4(–9) cm × 1–1.2(–1.5) mm, (replum not constricted between seeds);

ovules 76–128 per ovary;

style usually clavate to subclavate, rarely cylindrical, (0.5–)0.8–2(–3) mm.

divaricate to ascending, torulose, straight or slightly curved, terete, (0.6–)1–2.5(–4.2) cm × 0.7–1(–1.8) mm;

ovules 22–50 per ovary;

style cylindrical, 0.5–1.5(–2.5) mm.

Seeds

0.7–1.3(–1.5) × 0.5–0.8 mm.

(flattened), 1–1.5(–1.7) × 0.5–0.7(–1) mm.

2n

= 26.

Thelypodium wrightii

Thelypodium crispum

Phenology Flowering (Mar-)Jun–Oct. Flowering Jun–Aug.
Habitat Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands Alkaline meadows and desert flats, mineralized ground near hot springs, desert shrub communities
Elevation 1200-2300 m (3900-7500 ft) 1200-3000(-3200) m (3900-9800(-10500) ft)
Distribution
from FNA
AZ; CO; NM; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Hidalgo)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
Discussion

Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp. oklahomensis and subsp. wrightii, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 738. FNA vol. 7, p. 730.
Parent taxa Brassicaceae > tribe Thelypodieae > Thelypodium Brassicaceae > tribe Thelypodieae > Thelypodium
Sibling taxa
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum
T. brachycarpum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
Synonyms Stanleyella wrightii, T. wrightii subsp. oklahomensis Thelypodiopsis crispa, T. brachycarpum var. crispum
Name authority A. Gray: Smithsonian Contr. Knowl. 3(5): 7. (1852) Greene ex Payson: Ann. Missouri Bot. Gard. 9: 264. (1923)
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