Thelypodium wrightii |
Thelypodium |
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Wright's stanleyella, Wright's thelypody |
thelypody |
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Habit | Biennials; slightly glaucous, glabrous. | Biennials, perennials, or, rarely, annuals; not scapose; glabrous or pubescent. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | branched distally, (6.5–)9.5–22.5(–28) dm. |
(simple or few to several from base), usually erect, rarely decumbent, branched basally and/or distally, (glabrous or pubescent). |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate or not, petiolate, blade margins usually entire, dentate, lyrate or pinnately lobed, rarely laciniate; cauline petiolate or sessile, blade (base cuneate, attenuate, auriculate, sagittate, or amplexicaul), margins often entire, sometimes dentate or pinnately lobed. |
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Basal leaves | (and proximal cauline): petiole (2–)4.5–8.5(–13.7) cm; blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5–)9.5–22.5(–28) cm × 28–55(–75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate). |
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Cauline leaves | petiolate; blade lanceolate to linear-lanceolate, 3.5–8.5(–13.5) cm × 7–14(–23) mm, margins usually entire or dentate, rarely lobed. |
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Racemes | somewhat lax, considerably elongated in fruit. |
(corymbose, dense or lax), usually slightly to considerably elongated in fruit (sometimes not elongated in T. integrifolium). |
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Flowers | sepals spreading or ascending oblong to linear, (3–)4–6(–7) × (0.8–)1–1.3(–1.5) mm; petals usually white, rarely lavender, linear or oblong, 4–7.5(–9) × 1–1.8(–2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base); nectar glands confluent; filaments equal, (3–)3.5–6.5(–8.5) mm; anthers (1–)1.5–2.5(–3) mm, circinately coiled; gynophore 0.2–2(–5) mm. |
sepals usually erect or ascending, rarely spreading to reflexed, ovate to oblong, linear, lanceolate, or oblanceolate, lateral pair slightly saccate or not basally; petals (erect or spreading), white, lavender, or purple, spatulate to obovate, or oblanceolate to linear, (margins crisped or not), claw differentiated or not from blade, (apex rounded); stamens subequal or tetradynamous, (exserted or included); filaments (erect or spreading, usually distinct, very rarely median ones united), not dilated basally; anthers usually linear to linear-oblong, rarely oblong or ovate, (sometimes apiculate, often circinately coiled after dehiscence); nectar glands confluent and subtending bases of stamens, or 2 or 4 and lateral. |
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Fruiting pedicels | horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5–)7–13(–17) mm, slightly flattened at base. |
usually horizontal, erect to ascending, or divaricate, rarely reflexed, slender or stout, (flattened or not basally, glabrous). |
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Fruits | horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5–)3.8–7.4(–9) cm × 1–1.2(–1.5) mm, (replum not constricted between seeds); ovules 76–128 per ovary; style usually clavate to subclavate, rarely cylindrical, (0.5–)0.8–2(–3) mm. |
stipitate, linear, torulose or smooth, terete, slightly 4-angled, or flattened; valves each with prominent midvein, glabrous; replum rounded; septum complete; ovules 12–128 per ovary; style distinct, (often cylindrical, rarely subclavate or subconical); stigma capitate, entire. |
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Seeds | 0.7–1.3(–1.5) × 0.5–0.8 mm. |
uniseriate, plump or flattened, not winged, usually oblong, rarely ovate; seed coat (minutely reticulate), not mucilaginous when wetted; cotyledons oblique, rarely incumbent or accumbent. |
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x | = 13. |
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Thelypodium wrightii |
Thelypodium |
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Phenology | Flowering (Mar-)Jun–Oct. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 1200-2300 m (3900-7500 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
AZ; CO; NM; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Hidalgo)
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w North America; n Mexico; wc North America |
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Discussion | Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp. oklahomensis and subsp. wrightii, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 16 (16 in the flora). As recognized herein and by recent authors (e.g., I. A. Al-Shehbaz 1973; R. C. Rollins 1993), Thelypodium is somewhat heterogeneous and the segregate Stanleyella might merit recognition, as by E. B. Payson (1923). Thelypodium has erect sepals, petals, and stamens, terete fruits, prominently veined fruit septa, and, often, cylindrical styles. In contrast, species of Stanleylla have spreading sepals, petals, and stamens, flattened fruits, veinless fruit septa, and clavate, subclavate, or subconical styles. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 738. | FNA vol. 7, p. 728. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Thelypodieae > Thelypodium | Brassicaceae > tribe Thelypodieae | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Stanleyella wrightii, T. wrightii subsp. oklahomensis | Pachypodium, Pleurophragma, Stanleyella | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | A. Gray: Smithsonian Contr. Knowl. 3(5): 7. (1852) | Endlicher: Gen. Pl. 11: 876. (1839) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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