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arrow thelypody, slender thelypody

entire leaf thelypodium, entire-leaf thelypody

Habit Biennials or perennials; (short-lived); sometimes glaucous, glabrous throughout or sparsely to densely pubescent. Biennials; somewhat glaucous throughout, glabrous.
Stems

branched often proximally and distally, (2–)3–8(–12.5) dm, (glabrous or pubescent).

paniculately branched distally, (2–)4.5–17(–28) dm.

Basal leaves

petiole (1–)2–8(–14) cm, ciliate;

blade usually oblanceolate to ovate, rarely oblong, (2–)6.5–20(–29) cm × 10–42(–50) mm, margins entire, (surfaces glabrous or pubescent).

petiole (0.5–)1.2–10(–15) cm;

blade usually oblong to lanceolate, or oblanceolate to spatulate, rarely ovate or obovate, (3.7–)5–31(–54) cm × (12–)16–78(–140) mm, margins usually entire, rarely dentate or repand.

Cauline leaves

(ascending);

sessile;

blade lanceolate to oblong-lanceolate, (0.8–)1.6–8.5(–14) cm × 2–13(–28) mm, (base amplexicaul to sagittate), margins entire, (surfaces glabrous or pubescent).

(ascending, not appressed to stem);

sessile;

blade linear to linear-lanceolate, (1–)2.2–8.3(–19.5) cm × (2–)3–11(–25) mm, (base cuneate to attenuate, not auriculate), margins entire.

Racemes

sometimes corymbose, lax or dense, elongated or not in fruit, (flower buds ovate to lanceolate).

dense, slightly or considerably elongated in fruit, (flower buds narrowly oblong).

Flowers

sepals erect, ovate to oblong, (2.5–)3–6(–10) × (1–)1.5–2(–2.5) mm;

petals white or lavender to purple, oblanceolate to spatulate or linear, (5–)7–14(–19) × (0.5–)1–3(–4) mm, margins slightly crisped between blade and claw, claw differentiated from blade, [slender, (2–)3–6.5(–10) mm, narrowest at base];

nectar glands confluent and subtending bases of stamens, sometimes only lateral;

filaments tetradynamous, median pairs 3–5.5(–8) mm, lateral pair (2–)2.5–4.5(–7) mm;

anthers included or partially exserted, oblong, (1.5–)2–3.2(–5) mm, not circinately coiled;

gynophore (stout), 0.5–1(–2) mm.

sepals erect, linear to linear-oblong, (3–)3.5–5.5(–7.5) × (0.5–)0.8–1(–1.5) mm;

petals white or lavender to purple, spatulate to oblanceolate, (4.5–)6–9(–13) × (0.5–)0.8–1.5(–2.3) mm, margins not crisped, claw differentiated from blade, [slender, (2–)3–4.5(–7) mm, narrowest at base];

nectar glands lateral, flat or toothlike, median glands absent;

filaments subequal to slightly tetradynamous, (3–)4.5–7.5(–12.5) mm;

anthers exserted, linear to narrowly oblong, (1–)1.5–2.5(–4.5) mm, not circinately coiled;

gynophore 0.5–2(–5.5) mm.

Fruiting pedicels

horizontal to divaricate, often curved upward, stout or slender, (2.5–)3.5–11(–20) mm, not flattened at base.

usually horizontal to divaricate, rarely ascending or reflexed, usually straight, rarely incurved, slender or stout, (2–)3–9(–13) mm, often strongly flattened, rarely not flattened at base.

Fruits

erect, torulose, straight or slightly incurved (forming arc with pedicel), usually terete, rarely 4-angled, 1–5.3(–6.9) cm × (0.5–)0.8–1(–1.2) mm;

ovules 20–62per ovary;

style cylindrical, 0.5–1.5(–3.3) mm.

usually horizontal to divaricate-ascending, or ascending, rarely reflexed, usually torulose, rarely submoniliform, usually strongly incurved, sometimes straight, rarely arcuate, terete, (0.8–)1.4–6.5(–8) cm × 1–1.3(–2) mm;

ovules 14–40 per ovary;

style usually cylindrical, rarely subclavate, 0.5–1.3(–3.2) mm.

Seeds

(flattened), (0.7–)1–1.3(–1.5) × 0.5–0.7(–1) mm.

(1–)1.2–2(–2.3) × 0.7–1(–1.3) mm.

Thelypodium sagittatum

Thelypodium integrifolium

Distribution
from FNA
CO; ID; MT; NV; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; ND; NE; NM; NV; OR; SD; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 5 (5 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Racemes often dense; fruiting pedicels 5-11(-20) mm; petals spatulate to oblanceolate, 1-3(-4) mm wide; median nectar glands often absent.
subsp. sagittatum
1. Racemes lax; fruiting pedicels (2.5-)3.5-7(-8) mm; petals linear to linear-oblanceolate, 0.5-1 (-1.5) mm wide; median nectar glands present.
subsp. ovalifolium
1. Petals white; fruiting pedicels whitish, stout, (5-) 6-11(-13) mm
→ 2
1. Petals usually lavender to purple, rarely white; fruiting pedicels not whitish, slender and 2-10 (-13) mm, or stout and 2-5(-6) mm
→ 3
2. Petals (5-)6-9(-10) mm; fruiting pedicels horizontal to divaricate, (5-)6-9(-13) mm; fruits (1.4-)1.9-3.7(-4.5) cm; gynophores (0.5-)1-3 mm; se California, s Nevada, sw Utah.
subsp. affine
2. Petals (7.5-)8-11(-13) mm; fruiting pedicels usually horizontal to reflexed, rarely divaricate, (6-)7-11(-13) mm; fruits (2.2-) 3.5-6.5(-8) cm; gynophores (1-)1.5-4 (-5.5) mm; Grand Canyon, Colorado River, Arizona.
subsp. longicarpum
3. Fruiting pedicels often slender, sometimes stout, not or slightly flattened at base, (4-)6- 10 (-13) mm.
subsp. integrifolium
3. Fruiting pedicels stout, strongly flattened at base, 2-5(-6) mm
→ 4
4. Racemes often strongly congested, central rachises (0.6-)1.2-7(-9) cm; gynophores stout, 0.5-1(-3) mm; e California, Nevada, s Oregon, nw Utah.
subsp. complanatum
4. Racemes somewhat congested, central rachises (3-)8-20(-28) cm; gynophores slender, (0.5-)0.8-3(-4) mm; ne Arizona, w Colorado, nw New Mexico, e Utah.
subsp. gracilipes
Source FNA vol. 7, p. 736. FNA vol. 7, p. 732.
Parent taxa Brassicaceae > tribe Thelypodieae > Thelypodium Brassicaceae > tribe Thelypodieae > Thelypodium
Sibling taxa
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. stenopetalum, T. texanum, T. wrightii
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
Subordinate taxa
T. sagittatum subsp. ovalifolium, T. sagittatum subsp. sagittatum
T. integrifolium subsp. affine, T. integrifolium subsp. complanatum, T. integrifolium subsp. gracilipes, T. integrifolium subsp. integrifolium, T. integrifolium subsp. longicarpum
Synonyms Streptanthus sagittatus, Pachypodium sagittatum, Thelypodiopsis nuttallii, Thelypodiopsis sagittata, Thelypodiopsis torulosa, T. nuttallii, T. torulosum Pachypodium integrifolium, Pleurophragma integrifolium
Name authority (Nuttall) Endlicher: in W. G. Walpers, Repert. Bot. Syst. 1: 172. (1842) (Nuttall) Endlicher: in W. G. Walpers, Repert. Bot. Syst. 1: 172. (1842)
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