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catleaf, droopflower thelypody, loose-flower thelypody

Howell's thelypody

Habit Biennials; somewhat glaucous, glabrous throughout or pubescent basally. Biennials or, rarely, perennials; (short-lived); glaucous, glabrous (except petioles).
Stems

branched distally, (1.5–)3–14(–23.5) dm, (glabrous throughout or sparsely to densely hirsute proximally).

branched distally, 1–8(–9.3) dm.

Basal leaves

(and proximal cauline): petiole (1–)1.7–8(–11) cm, pubescent or glabrous;

blade oblanceolate in outline (lateral lobes usually oblong to linear, rarely deltate), (4–)7.2–20.5(–30) cm × (6–)10–40(–100) mm, margins pinnately lobed or lyrate (lobes entire or dentate).

petiole 0.5–2 cm, ciliate;

blade often oblanceolate to spatulate, sometimes oblong or lanceolate, 2–10(–13.5) cm × 10–23(–40) mm, margins often lyrate to dentate, sometimes entire or repand.

Cauline leaves

petiolate;

blade usually lanceolate to linear, sometimes oblanceolate to oblong, 1.5–7(–10) cm × 3–12(–20) mm, margins usually entire or repand, rarely lobed.

(ascending);

sessile;

blade often lanceolate to linear, sometimes oblong, (0.8–)1–9.7(–14.5) cm × 2–10(–30) mm, (base usually amplexicaul to strongly sagittate, rarely auriculate), margins entire.

Racemes

corymbose, somewhat lax, considerably elongated in fruit.

lax, elongated in fruit, (flower buds ovate to lanceolate).

Flowers

sepals (equal), ascending, oblong, (2.5–)3–5(–5.5) × 1–1.7(–3) mm, (lateral pair not saccate);

petals usually white, rarely lavender, usually spatulate, rarely obovate, 5–7.5(–9.5) × 1.5–2.5(–3.5) mm, margins not crisped, claw obscurely differentiated from blade (widest at base);

nectar glands continuous, flat, subtending bases of stamens;

filaments tetradynamous, median pairs 3–4.5(–5.5) mm, lateral pair 2.5–3.5(–4.5) mm;

anthers included, oblong, (1–)1.5–2.2(–2.5) mm, not circinately coiled;

gynophore 0.5–0.8(–1.5) mm.

sepals erect, linear or lanceolate, (3.5–)5–8.5(–11.5) × (0.7–)1–2(–2.5) mm;

petals lavender to purple, linear or oblanceolate to spatulate, (6–)9–16.5(–22) × 0.5–2.7(–3) mm, margins partially crisped, claw differentiated from blade, [slender, (2.5–)4–7(–11) mm, narrowest at base];

nectar glands confluent, subtending bases of stamens;

filaments slightly tetradynamous, median pairs (3–)5.5–9(–12) mm, lateral pair (2.5–)4.5–8(–10) mm;

anthers slightly exserted to included, linear, (1.5–)2–4(–5.5) mm, not circinately coiled;

gynophore (stout), 0.5–1(–3.5) mm.

Fruiting pedicels

often divaricate, sometimes reflexed, rarely horizontal or divaricate-ascending, straight, slender, (4–)5–13(–19) mm, somewhat flattened at base.

usually ascending, rarely divaricate-ascending, usually straight, rarely slightly incurved, stout, (2.5–)3.5–8(–14.5) mm, not flattened at base.

Fruits

divaricate to reflexed, submoniliform to strongly torulose, straight or curved, terete, (2–)3–6.6(–7.4) cm × 0.7–1(–1.5) mm, (replum constricted between seeds);

ovules 26–62 per ovary;

style usually clavate to subclavate, rarely cylindrical, (0.5–)0.8–2(–5) mm.

often ascending, sometimes erect or divaricate, slightly torulose, straight or slightly incurved (forming straight line with pedicel), terete, 1.5–6.5(–8.2) cm × 1–1.3(–2) mm;

ovules 22–40 per ovary;

style cylindrical or subclavate, (0.5–)1–2.7(–4.3) mm.

Seeds

(oblong), (1–)1.3–1.8(–2) × 0.5–1 mm.

(1–)1.2–1.7(–2) × 0.5–0.7 mm.

Thelypodium laxiflorum

Thelypodium howellii

Phenology Flowering May–Sep.
Habitat Talus, rocky slopes, cliffs, pinyon-juniper-brush communities
Elevation 1500-3100 m (4900-10200 ft)
Distribution
from FNA
CO; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Streptanthus howellii (S. Watson) M. E. Jones is a later homonym of S. howellii S. Watson 1885.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals usually linear to narrowly oblanceolate, rarely spatulate, 0.5-1.2(-3) mm wide; median filaments connate most of their lengths; ne California, c, s Oregon, s Washington.
subsp. howellii
1. Petals usually spatulate, rarely oblanceolate, (1.2-) 1.5-2.7(-3) mm wide; median filaments distinct; e Oregon.
subsp. spectabilis
Source FNA vol. 7, p. 735. FNA vol. 7, p. 732.
Parent taxa Brassicaceae > tribe Thelypodieae > Thelypodium Brassicaceae > tribe Thelypodieae > Thelypodium
Sibling taxa
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laciniatum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
Subordinate taxa
T. howellii subsp. howellii, T. howellii subsp. spectabilis
Synonyms Stanleyella wrightii var. tenella, T. wrightii var. tenellum Cartiera howellii, Thelypodiopsis howellii
Name authority Al-Shehbaz: Contr. Gray Herb. 204: 129, plate 22A. (1973) S. Watson: Proc. Amer. Acad. Arts 21: 445. (1886)
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