Thelypodium laciniatum |
Thelypodium |
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cut-leaf thelypody, feathery thelypodium, many-flower thelypody, thick-leaf thelypody, thick-leaf thelypody (var. laciniatum), western thelypodium |
thelypody |
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Habit | Biennials; somewhat glaucous, glabrous throughout. | Biennials, perennials, or, rarely, annuals; not scapose; glabrous or pubescent. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | branched proximally and/or distally, (1.3–)2.6–10(–14.5) dm. |
(simple or few to several from base), usually erect, rarely decumbent, branched basally and/or distally, (glabrous or pubescent). |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate or not, petiolate, blade margins usually entire, dentate, lyrate or pinnately lobed, rarely laciniate; cauline petiolate or sessile, blade (base cuneate, attenuate, auriculate, sagittate, or amplexicaul), margins often entire, sometimes dentate or pinnately lobed. |
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Basal leaves | (and proximal cauline): petiole (1–)1.5–10(–15) cm; blade lanceolate to deltate-lanceolate, or oblong to ovate (lateral lobes oblong to ovate), (4.3–)6.8–24(–45) cm × (10–)18–85(–140) mm, margins pinnately lobed or laciniate (lobes sinuate, laciniate, or dentate). |
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Cauline leaves | petiolate; blade lanceolate to linear-lanceolate, similar to basal, smaller distally, margins entire or sinuate to laciniate. |
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Racemes | dense, slightly elongated in fruit (to 9.5 dm, flower buds narrowly oblong). |
(corymbose, dense or lax), usually slightly to considerably elongated in fruit (sometimes not elongated in T. integrifolium). |
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Flowers | sepals erect, oblong to linear-oblong, (3.5–)4–7.5(–9.5) × (0.8–)1–2(–2.5) mm; petals white or purple, often linear, sometimes linear-lanceolate, (6–)8.5–18(–20) × 0.3–0.8(–1.5) mm, margins slightly crisped, claw strongly differentiated from blade, [(2.5–)3–6(–7) mm, widest at base]; nectar glands confluent, lateral and median; filaments subequal, (4.5–)6.5–12(–15) mm; anthers exserted, linear, 2–4(–5.5) mm, circinately coiled, (apiculate); gynophore (0.5–)1–5(–8.5) mm. |
sepals usually erect or ascending, rarely spreading to reflexed, ovate to oblong, linear, lanceolate, or oblanceolate, lateral pair slightly saccate or not basally; petals (erect or spreading), white, lavender, or purple, spatulate to obovate, or oblanceolate to linear, (margins crisped or not), claw differentiated or not from blade, (apex rounded); stamens subequal or tetradynamous, (exserted or included); filaments (erect or spreading, usually distinct, very rarely median ones united), not dilated basally; anthers usually linear to linear-oblong, rarely oblong or ovate, (sometimes apiculate, often circinately coiled after dehiscence); nectar glands confluent and subtending bases of stamens, or 2 or 4 and lateral. |
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Fruiting pedicels | horizontal to, rarely, divaricate, straight, often stout, (2–)3–6.5(–15) mm, often flattened at base. |
usually horizontal, erect to ascending, or divaricate, rarely reflexed, slender or stout, (flattened or not basally, glabrous). |
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Fruits | divaricate-ascending to slightly reflexed, torulose, somewhat tortuous, terete, (2.5–)3.5–10(–12) cm × (0.7–)1–1.5(–2) mm; ovules 56–108 per ovary; style usually cylindrical, rarely subclavate, (0.3–)0.7–2.5(–4) mm. |
stipitate, linear, torulose or smooth, terete, slightly 4-angled, or flattened; valves each with prominent midvein, glabrous; replum rounded; septum complete; ovules 12–128 per ovary; style distinct, (often cylindrical, rarely subclavate or subconical); stigma capitate, entire. |
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Seeds | (0.7–)1–1.5(–1.8) × 0.5–0.7(–1) mm. |
uniseriate, plump or flattened, not winged, usually oblong, rarely ovate; seed coat (minutely reticulate), not mucilaginous when wetted; cotyledons oblique, rarely incumbent or accumbent. |
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x | = 13. |
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2n | = 26. |
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Thelypodium laciniatum |
Thelypodium |
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Phenology | Flowering Apr–Aug. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Rock crevices, cliffs, rocky outcrops, among boulders, serpentine rock, talus, canyon walls, limestone ledges | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 0-2400 m (0-7900 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA; ID; NV; OR; WA; BC
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w North America; n Mexico; wc North America |
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Discussion | Species 16 (16 in the flora). As recognized herein and by recent authors (e.g., I. A. Al-Shehbaz 1973; R. C. Rollins 1993), Thelypodium is somewhat heterogeneous and the segregate Stanleyella might merit recognition, as by E. B. Payson (1923). Thelypodium has erect sepals, petals, and stamens, terete fruits, prominently veined fruit septa, and, often, cylindrical styles. In contrast, species of Stanleylla have spreading sepals, petals, and stamens, flattened fruits, veinless fruit septa, and clavate, subclavate, or subconical styles. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 734. | FNA vol. 7, p. 728. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Thelypodieae > Thelypodium | Brassicaceae > tribe Thelypodieae | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Synonyms | Macropodium laciniatum, Pachypodium laciniatum, T. laciniatum var. streptanthoides, T. leptosepalum, T. streptanthoides | Pachypodium, Pleurophragma, Stanleyella | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | (Hooker) Endlicher: in W. G. Walpers, Repert. Bot. Syst. 1: 172. (1842) | Endlicher: Gen. Pl. 11: 876. (1839) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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