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cut-leaf thelypody, feathery thelypodium, many-flower thelypody, thick-leaf thelypody, thick-leaf thelypody (var. laciniatum), western thelypodium

Howell's thelypody

Habit Biennials; somewhat glaucous, glabrous throughout. Biennials or, rarely, perennials; (short-lived); glaucous, glabrous (except petioles).
Stems

branched proximally and/or distally, (1.3–)2.6–10(–14.5) dm.

branched distally, 1–8(–9.3) dm.

Basal leaves

(and proximal cauline): petiole (1–)1.5–10(–15) cm;

blade lanceolate to deltate-lanceolate, or oblong to ovate (lateral lobes oblong to ovate), (4.3–)6.8–24(–45) cm × (10–)18–85(–140) mm, margins pinnately lobed or laciniate (lobes sinuate, laciniate, or dentate).

petiole 0.5–2 cm, ciliate;

blade often oblanceolate to spatulate, sometimes oblong or lanceolate, 2–10(–13.5) cm × 10–23(–40) mm, margins often lyrate to dentate, sometimes entire or repand.

Cauline leaves

petiolate;

blade lanceolate to linear-lanceolate, similar to basal, smaller distally, margins entire or sinuate to laciniate.

(ascending);

sessile;

blade often lanceolate to linear, sometimes oblong, (0.8–)1–9.7(–14.5) cm × 2–10(–30) mm, (base usually amplexicaul to strongly sagittate, rarely auriculate), margins entire.

Racemes

dense, slightly elongated in fruit (to 9.5 dm, flower buds narrowly oblong).

lax, elongated in fruit, (flower buds ovate to lanceolate).

Flowers

sepals erect, oblong to linear-oblong, (3.5–)4–7.5(–9.5) × (0.8–)1–2(–2.5) mm;

petals white or purple, often linear, sometimes linear-lanceolate, (6–)8.5–18(–20) × 0.3–0.8(–1.5) mm, margins slightly crisped, claw strongly differentiated from blade, [(2.5–)3–6(–7) mm, widest at base];

nectar glands confluent, lateral and median;

filaments subequal, (4.5–)6.5–12(–15) mm;

anthers exserted, linear, 2–4(–5.5) mm, circinately coiled, (apiculate);

gynophore (0.5–)1–5(–8.5) mm.

sepals erect, linear or lanceolate, (3.5–)5–8.5(–11.5) × (0.7–)1–2(–2.5) mm;

petals lavender to purple, linear or oblanceolate to spatulate, (6–)9–16.5(–22) × 0.5–2.7(–3) mm, margins partially crisped, claw differentiated from blade, [slender, (2.5–)4–7(–11) mm, narrowest at base];

nectar glands confluent, subtending bases of stamens;

filaments slightly tetradynamous, median pairs (3–)5.5–9(–12) mm, lateral pair (2.5–)4.5–8(–10) mm;

anthers slightly exserted to included, linear, (1.5–)2–4(–5.5) mm, not circinately coiled;

gynophore (stout), 0.5–1(–3.5) mm.

Fruiting pedicels

horizontal to, rarely, divaricate, straight, often stout, (2–)3–6.5(–15) mm, often flattened at base.

usually ascending, rarely divaricate-ascending, usually straight, rarely slightly incurved, stout, (2.5–)3.5–8(–14.5) mm, not flattened at base.

Fruits

divaricate-ascending to slightly reflexed, torulose, somewhat tortuous, terete, (2.5–)3.5–10(–12) cm × (0.7–)1–1.5(–2) mm;

ovules 56–108 per ovary;

style usually cylindrical, rarely subclavate, (0.3–)0.7–2.5(–4) mm.

often ascending, sometimes erect or divaricate, slightly torulose, straight or slightly incurved (forming straight line with pedicel), terete, 1.5–6.5(–8.2) cm × 1–1.3(–2) mm;

ovules 22–40 per ovary;

style cylindrical or subclavate, (0.5–)1–2.7(–4.3) mm.

Seeds

(0.7–)1–1.5(–1.8) × 0.5–0.7(–1) mm.

(1–)1.2–1.7(–2) × 0.5–0.7 mm.

2n

= 26.

Thelypodium laciniatum

Thelypodium howellii

Phenology Flowering Apr–Aug.
Habitat Rock crevices, cliffs, rocky outcrops, among boulders, serpentine rock, talus, canyon walls, limestone ledges
Elevation 0-2400 m (0-7900 ft)
Distribution
from FNA
CA; ID; NV; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Streptanthus howellii (S. Watson) M. E. Jones is a later homonym of S. howellii S. Watson 1885.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals usually linear to narrowly oblanceolate, rarely spatulate, 0.5-1.2(-3) mm wide; median filaments connate most of their lengths; ne California, c, s Oregon, s Washington.
subsp. howellii
1. Petals usually spatulate, rarely oblanceolate, (1.2-) 1.5-2.7(-3) mm wide; median filaments distinct; e Oregon.
subsp. spectabilis
Source FNA vol. 7, p. 734. FNA vol. 7, p. 732.
Parent taxa Brassicaceae > tribe Thelypodieae > Thelypodium Brassicaceae > tribe Thelypodieae > Thelypodium
Sibling taxa
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
Subordinate taxa
T. howellii subsp. howellii, T. howellii subsp. spectabilis
Synonyms Macropodium laciniatum, Pachypodium laciniatum, T. laciniatum var. streptanthoides, T. leptosepalum, T. streptanthoides Cartiera howellii, Thelypodiopsis howellii
Name authority (Hooker) Endlicher: in W. G. Walpers, Repert. Bot. Syst. 1: 172. (1842) S. Watson: Proc. Amer. Acad. Arts 21: 445. (1886)
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