Thelypodium laciniatum |
Thelypodium howellii |
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cut-leaf thelypody, feathery thelypodium, many-flower thelypody, thick-leaf thelypody, thick-leaf thelypody (var. laciniatum), western thelypodium |
Howell's thelypody |
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Habit | Biennials; somewhat glaucous, glabrous throughout. | Biennials or, rarely, perennials; (short-lived); glaucous, glabrous (except petioles). | ||||
Stems | branched proximally and/or distally, (1.3–)2.6–10(–14.5) dm. |
branched distally, 1–8(–9.3) dm. |
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Basal leaves | (and proximal cauline): petiole (1–)1.5–10(–15) cm; blade lanceolate to deltate-lanceolate, or oblong to ovate (lateral lobes oblong to ovate), (4.3–)6.8–24(–45) cm × (10–)18–85(–140) mm, margins pinnately lobed or laciniate (lobes sinuate, laciniate, or dentate). |
petiole 0.5–2 cm, ciliate; blade often oblanceolate to spatulate, sometimes oblong or lanceolate, 2–10(–13.5) cm × 10–23(–40) mm, margins often lyrate to dentate, sometimes entire or repand. |
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Cauline leaves | petiolate; blade lanceolate to linear-lanceolate, similar to basal, smaller distally, margins entire or sinuate to laciniate. |
(ascending); sessile; blade often lanceolate to linear, sometimes oblong, (0.8–)1–9.7(–14.5) cm × 2–10(–30) mm, (base usually amplexicaul to strongly sagittate, rarely auriculate), margins entire. |
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Racemes | dense, slightly elongated in fruit (to 9.5 dm, flower buds narrowly oblong). |
lax, elongated in fruit, (flower buds ovate to lanceolate). |
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Flowers | sepals erect, oblong to linear-oblong, (3.5–)4–7.5(–9.5) × (0.8–)1–2(–2.5) mm; petals white or purple, often linear, sometimes linear-lanceolate, (6–)8.5–18(–20) × 0.3–0.8(–1.5) mm, margins slightly crisped, claw strongly differentiated from blade, [(2.5–)3–6(–7) mm, widest at base]; nectar glands confluent, lateral and median; filaments subequal, (4.5–)6.5–12(–15) mm; anthers exserted, linear, 2–4(–5.5) mm, circinately coiled, (apiculate); gynophore (0.5–)1–5(–8.5) mm. |
sepals erect, linear or lanceolate, (3.5–)5–8.5(–11.5) × (0.7–)1–2(–2.5) mm; petals lavender to purple, linear or oblanceolate to spatulate, (6–)9–16.5(–22) × 0.5–2.7(–3) mm, margins partially crisped, claw differentiated from blade, [slender, (2.5–)4–7(–11) mm, narrowest at base]; nectar glands confluent, subtending bases of stamens; filaments slightly tetradynamous, median pairs (3–)5.5–9(–12) mm, lateral pair (2.5–)4.5–8(–10) mm; anthers slightly exserted to included, linear, (1.5–)2–4(–5.5) mm, not circinately coiled; gynophore (stout), 0.5–1(–3.5) mm. |
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Fruiting pedicels | horizontal to, rarely, divaricate, straight, often stout, (2–)3–6.5(–15) mm, often flattened at base. |
usually ascending, rarely divaricate-ascending, usually straight, rarely slightly incurved, stout, (2.5–)3.5–8(–14.5) mm, not flattened at base. |
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Fruits | divaricate-ascending to slightly reflexed, torulose, somewhat tortuous, terete, (2.5–)3.5–10(–12) cm × (0.7–)1–1.5(–2) mm; ovules 56–108 per ovary; style usually cylindrical, rarely subclavate, (0.3–)0.7–2.5(–4) mm. |
often ascending, sometimes erect or divaricate, slightly torulose, straight or slightly incurved (forming straight line with pedicel), terete, 1.5–6.5(–8.2) cm × 1–1.3(–2) mm; ovules 22–40 per ovary; style cylindrical or subclavate, (0.5–)1–2.7(–4.3) mm. |
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Seeds | (0.7–)1–1.5(–1.8) × 0.5–0.7(–1) mm. |
(1–)1.2–1.7(–2) × 0.5–0.7 mm. |
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2n | = 26. |
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Thelypodium laciniatum |
Thelypodium howellii |
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Phenology | Flowering Apr–Aug. | |||||
Habitat | Rock crevices, cliffs, rocky outcrops, among boulders, serpentine rock, talus, canyon walls, limestone ledges | |||||
Elevation | 0-2400 m (0-7900 ft) | |||||
Distribution |
CA; ID; NV; OR; WA; BC
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CA; OR; WA
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Discussion | Subspecies 2 (2 in the flora). Streptanthus howellii (S. Watson) M. E. Jones is a later homonym of S. howellii S. Watson 1885. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 734. | FNA vol. 7, p. 732. | ||||
Parent taxa | Brassicaceae > tribe Thelypodieae > Thelypodium | Brassicaceae > tribe Thelypodieae > Thelypodium | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Macropodium laciniatum, Pachypodium laciniatum, T. laciniatum var. streptanthoides, T. leptosepalum, T. streptanthoides | Cartiera howellii, Thelypodiopsis howellii | ||||
Name authority | (Hooker) Endlicher: in W. G. Walpers, Repert. Bot. Syst. 1: 172. (1842) | S. Watson: Proc. Amer. Acad. Arts 21: 445. (1886) | ||||
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