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entire leaf thelypodium, entire-leaf thelypody

northwestern thelypody

Habit Biennials; somewhat glaucous throughout, glabrous. Biennials or perennials; (short-lived); glaucous, glabrous or sparsely pubescent.
Stems

paniculately branched distally, (2–)4.5–17(–28) dm.

branched often proximally and distally, (1.4–)2–6.5(–7.5) dm, (glabrous or pubescent).

Basal leaves

petiole (0.5–)1.2–10(–15) cm;

blade usually oblong to lanceolate, or oblanceolate to spatulate, rarely ovate or obovate, (3.7–)5–31(–54) cm × (12–)16–78(–140) mm, margins usually entire, rarely dentate or repand.

petiole (0.8–)2–4(–6.5) cm, ciliate;

blade usually oblanceolate, rarely spatulate or oblong, (3–)6–15(–22) cm × (6–)10–25(–40) mm, margins entire, (surfaces glabrous).

Cauline leaves

(ascending, not appressed to stem);

sessile;

blade linear to linear-lanceolate, (1–)2.2–8.3(–19.5) cm × (2–)3–11(–25) mm, (base cuneate to attenuate, not auriculate), margins entire.

(ascending);

sessile;

blade lanceolate to linear-lanceolate, (1.7–)2.2–6(–9) cm × (3–)5–15(–30) mm, (base sagittate to somewhat amplexicaul), margins entire, (surfaces glabrous).

Racemes

dense, slightly or considerably elongated in fruit, (flower buds narrowly oblong).

corymbose, dense, terminal not elongated in fruit, (flower buds ovate).

Flowers

sepals erect, linear to linear-oblong, (3–)3.5–5.5(–7.5) × (0.5–)0.8–1(–1.5) mm;

petals white or lavender to purple, spatulate to oblanceolate, (4.5–)6–9(–13) × (0.5–)0.8–1.5(–2.3) mm, margins not crisped, claw differentiated from blade, [slender, (2–)3–4.5(–7) mm, narrowest at base];

nectar glands lateral, flat or toothlike, median glands absent;

filaments subequal to slightly tetradynamous, (3–)4.5–7.5(–12.5) mm;

anthers exserted, linear to narrowly oblong, (1–)1.5–2.5(–4.5) mm, not circinately coiled;

gynophore 0.5–2(–5.5) mm.

sepals erect, ovate to oblong-ovate, (3–)3.5–5(–6) × (1–)1.5–2 mm;

petals lavender to purple, spatulate to broadly obovate, (6.5–)8–11.5(–14) × 2.5–5(–6) mm, margins not crisped, claw strongly differentiated from blade, [slender, (2–)2.5–4(–5) mm, narrowest at base];

nectar glands confluent, subtending bases of stamens;

filaments tetradynamous, median pairs (3–)3.5–5(–6) mm, lateral pair (2–)2.5–4(–5) mm;

anthers included, ovate to ovate-oblong, 1–2 mm, not circinately coiled;

gynophore (stout), 0.5–0.8(–1.2) mm.

Fruiting pedicels

usually horizontal to divaricate, rarely ascending or reflexed, usually straight, rarely incurved, slender or stout, (2–)3–9(–13) mm, often strongly flattened, rarely not flattened at base.

usually horizontal to divaricate, rarely divaricate-ascending, often curved upward, slender, 6–11.5(–17) mm, slightly flattened at base.

Fruits

usually horizontal to divaricate-ascending, or ascending, rarely reflexed, usually torulose, rarely submoniliform, usually strongly incurved, sometimes straight, rarely arcuate, terete, (0.8–)1.4–6.5(–8) cm × 1–1.3(–2) mm;

ovules 14–40 per ovary;

style usually cylindrical, rarely subclavate, 0.5–1.3(–3.2) mm.

erect, slightly torulose, usually slightly incurved, rarely straight, terete, (1–)1.3–3.2(–5) cm × (0.8–)1.5–2.3 mm;

ovules 20–30 per ovary;

style cylindrical, (0.5–)1.5–1.7(–3) mm.

Seeds

(1–)1.2–2(–2.3) × 0.7–1(–1.3) mm.

(plump), (1.3–)1.5–2 × 0.8–1 mm.

Thelypodium integrifolium

Thelypodium paniculatum

Phenology Flowering Jun–Jul.
Habitat Wet sedge meadows, streamsides
Elevation 1800-2800 m (5900-9200 ft)
Distribution
from FNA
AZ; CA; CO; ID; MT; ND; NE; NM; NV; OR; SD; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ID; MT; WY
[BONAP county map]
Discussion

Subspecies 5 (5 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals white; fruiting pedicels whitish, stout, (5-) 6-11(-13) mm
→ 2
1. Petals usually lavender to purple, rarely white; fruiting pedicels not whitish, slender and 2-10 (-13) mm, or stout and 2-5(-6) mm
→ 3
2. Petals (5-)6-9(-10) mm; fruiting pedicels horizontal to divaricate, (5-)6-9(-13) mm; fruits (1.4-)1.9-3.7(-4.5) cm; gynophores (0.5-)1-3 mm; se California, s Nevada, sw Utah.
subsp. affine
2. Petals (7.5-)8-11(-13) mm; fruiting pedicels usually horizontal to reflexed, rarely divaricate, (6-)7-11(-13) mm; fruits (2.2-) 3.5-6.5(-8) cm; gynophores (1-)1.5-4 (-5.5) mm; Grand Canyon, Colorado River, Arizona.
subsp. longicarpum
3. Fruiting pedicels often slender, sometimes stout, not or slightly flattened at base, (4-)6- 10 (-13) mm.
subsp. integrifolium
3. Fruiting pedicels stout, strongly flattened at base, 2-5(-6) mm
→ 4
4. Racemes often strongly congested, central rachises (0.6-)1.2-7(-9) cm; gynophores stout, 0.5-1(-3) mm; e California, Nevada, s Oregon, nw Utah.
subsp. complanatum
4. Racemes somewhat congested, central rachises (3-)8-20(-28) cm; gynophores slender, (0.5-)0.8-3(-4) mm; ne Arizona, w Colorado, nw New Mexico, e Utah.
subsp. gracilipes
Source FNA vol. 7, p. 732. FNA vol. 7, p. 735.
Parent taxa Brassicaceae > tribe Thelypodieae > Thelypodium Brassicaceae > tribe Thelypodieae > Thelypodium
Sibling taxa
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
Subordinate taxa
T. integrifolium subsp. affine, T. integrifolium subsp. complanatum, T. integrifolium subsp. gracilipes, T. integrifolium subsp. integrifolium, T. integrifolium subsp. longicarpum
Synonyms Pachypodium integrifolium, Pleurophragma integrifolium Thelypodiopsis paniculata, T. sagittatum var. crassicarpum
Name authority (Nuttall) Endlicher: in W. G. Walpers, Repert. Bot. Syst. 1: 172. (1842) A. Nelson: Bull. Torrey Bot. Club 26: 126. (1899)
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