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entireleaf thelypody

entire leaf thelypodium, entire-leaf thelypody

Habit Biennials; somewhat glaucous throughout, glabrous.
Stems

paniculately branched distally, (2–)4.5–17(–28) dm.

Basal leaves

petiole (0.5–)1.2–10(–15) cm;

blade usually oblong to lanceolate, or oblanceolate to spatulate, rarely ovate or obovate, (3.7–)5–31(–54) cm × (12–)16–78(–140) mm, margins usually entire, rarely dentate or repand.

Cauline leaves

(ascending, not appressed to stem);

sessile;

blade linear to linear-lanceolate, (1–)2.2–8.3(–19.5) cm × (2–)3–11(–25) mm, (base cuneate to attenuate, not auriculate), margins entire.

Racemes

(moderately to strongly congested), not elongated in fruit;

central rachis (4.3–)8–20 cm.

dense, slightly or considerably elongated in fruit, (flower buds narrowly oblong).

Flowers

petals white, (7.5–)8–11(–13) mm;

gynophore stout, (1–)1.5–4(–5.5) mm.

sepals erect, linear to linear-oblong, (3–)3.5–5.5(–7.5) × (0.5–)0.8–1(–1.5) mm;

petals white or lavender to purple, spatulate to oblanceolate, (4.5–)6–9(–13) × (0.5–)0.8–1.5(–2.3) mm, margins not crisped, claw differentiated from blade, [slender, (2–)3–4.5(–7) mm, narrowest at base];

nectar glands lateral, flat or toothlike, median glands absent;

filaments subequal to slightly tetradynamous, (3–)4.5–7.5(–12.5) mm;

anthers exserted, linear to narrowly oblong, (1–)1.5–2.5(–4.5) mm, not circinately coiled;

gynophore 0.5–2(–5.5) mm.

Fruiting pedicels

usually horizontal to reflexed, rarely divaricate, straight, whitish, stout, (6–)7–11(–13) mm, strongly flattened at base.

usually horizontal to divaricate, rarely ascending or reflexed, usually straight, rarely incurved, slender or stout, (2–)3–9(–13) mm, often strongly flattened, rarely not flattened at base.

Fruits

usually horizontal to reflexed, rarely divaricate, straight or incurved to arcuate, (2.2–)3.5–6.5(–8) cm.

usually horizontal to divaricate-ascending, or ascending, rarely reflexed, usually torulose, rarely submoniliform, usually strongly incurved, sometimes straight, rarely arcuate, terete, (0.8–)1.4–6.5(–8) cm × 1–1.3(–2) mm;

ovules 14–40 per ovary;

style usually cylindrical, rarely subclavate, 0.5–1.3(–3.2) mm.

Seeds

(1–)1.2–2(–2.3) × 0.7–1(–1.3) mm.

Thelypodium integrifolium subsp. longicarpum

Thelypodium integrifolium

Phenology Flowering Aug–Oct.
Habitat Canyons, shrub communities
Elevation 500-1300 m (1600-4300 ft)
Distribution
from FNA
AZ
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; ND; NE; NM; NV; OR; SD; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Subspecies longicarpum is known from the Grand Canyon National Park and adjacent canyons of the Colorado River.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 5 (5 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals white; fruiting pedicels whitish, stout, (5-) 6-11(-13) mm
→ 2
1. Petals usually lavender to purple, rarely white; fruiting pedicels not whitish, slender and 2-10 (-13) mm, or stout and 2-5(-6) mm
→ 3
2. Petals (5-)6-9(-10) mm; fruiting pedicels horizontal to divaricate, (5-)6-9(-13) mm; fruits (1.4-)1.9-3.7(-4.5) cm; gynophores (0.5-)1-3 mm; se California, s Nevada, sw Utah.
subsp. affine
2. Petals (7.5-)8-11(-13) mm; fruiting pedicels usually horizontal to reflexed, rarely divaricate, (6-)7-11(-13) mm; fruits (2.2-) 3.5-6.5(-8) cm; gynophores (1-)1.5-4 (-5.5) mm; Grand Canyon, Colorado River, Arizona.
subsp. longicarpum
3. Fruiting pedicels often slender, sometimes stout, not or slightly flattened at base, (4-)6- 10 (-13) mm.
subsp. integrifolium
3. Fruiting pedicels stout, strongly flattened at base, 2-5(-6) mm
→ 4
4. Racemes often strongly congested, central rachises (0.6-)1.2-7(-9) cm; gynophores stout, 0.5-1(-3) mm; e California, Nevada, s Oregon, nw Utah.
subsp. complanatum
4. Racemes somewhat congested, central rachises (3-)8-20(-28) cm; gynophores slender, (0.5-)0.8-3(-4) mm; ne Arizona, w Colorado, nw New Mexico, e Utah.
subsp. gracilipes
Source FNA vol. 7, p. 734. FNA vol. 7, p. 732.
Parent taxa Brassicaceae > tribe Thelypodieae > Thelypodium > Thelypodium integrifolium Brassicaceae > tribe Thelypodieae > Thelypodium
Sibling taxa
T. integrifolium subsp. affine, T. integrifolium subsp. complanatum, T. integrifolium subsp. gracilipes, T. integrifolium subsp. integrifolium
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
Subordinate taxa
T. integrifolium subsp. affine, T. integrifolium subsp. complanatum, T. integrifolium subsp. gracilipes, T. integrifolium subsp. integrifolium, T. integrifolium subsp. longicarpum
Synonyms T. integrifolium var. longicarpum Pachypodium integrifolium, Pleurophragma integrifolium
Name authority Al-Shehbaz: Contr. Gray Herb. 204: 111. (1973) (Nuttall) Endlicher: in W. G. Walpers, Repert. Bot. Syst. 1: 172. (1842)
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