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white entire leaf thelypodium

entire leaf thelypodium, entire-leaf thelypody

Habit Biennials; somewhat glaucous throughout, glabrous.
Stems

paniculately branched distally, (2–)4.5–17(–28) dm.

Basal leaves

petiole (0.5–)1.2–10(–15) cm;

blade usually oblong to lanceolate, or oblanceolate to spatulate, rarely ovate or obovate, (3.7–)5–31(–54) cm × (12–)16–78(–140) mm, margins usually entire, rarely dentate or repand.

Cauline leaves

(ascending, not appressed to stem);

sessile;

blade linear to linear-lanceolate, (1–)2.2–8.3(–19.5) cm × (2–)3–11(–25) mm, (base cuneate to attenuate, not auriculate), margins entire.

Racemes

(congested), elongated in fruit;

central rachis (3.7–)4.5–12(–16.5) cm.

dense, slightly or considerably elongated in fruit, (flower buds narrowly oblong).

Flowers

petals white, (5–)6–9(–10) mm;

gynophore stout, (0.5–)1–3 mm.

sepals erect, linear to linear-oblong, (3–)3.5–5.5(–7.5) × (0.5–)0.8–1(–1.5) mm;

petals white or lavender to purple, spatulate to oblanceolate, (4.5–)6–9(–13) × (0.5–)0.8–1.5(–2.3) mm, margins not crisped, claw differentiated from blade, [slender, (2–)3–4.5(–7) mm, narrowest at base];

nectar glands lateral, flat or toothlike, median glands absent;

filaments subequal to slightly tetradynamous, (3–)4.5–7.5(–12.5) mm;

anthers exserted, linear to narrowly oblong, (1–)1.5–2.5(–4.5) mm, not circinately coiled;

gynophore 0.5–2(–5.5) mm.

Fruiting pedicels

horizontal to divaricate, straight, whitish, stout, (5–)6–9(–13) mm, moderately flattened at base.

usually horizontal to divaricate, rarely ascending or reflexed, usually straight, rarely incurved, slender or stout, (2–)3–9(–13) mm, often strongly flattened, rarely not flattened at base.

Fruits

usually horizontal or divaricate, rarely divaricate-ascending, incurved, (1.4–)1.9–3.7(–4.5) cm.

usually horizontal to divaricate-ascending, or ascending, rarely reflexed, usually torulose, rarely submoniliform, usually strongly incurved, sometimes straight, rarely arcuate, terete, (0.8–)1.4–6.5(–8) cm × 1–1.3(–2) mm;

ovules 14–40 per ovary;

style usually cylindrical, rarely subclavate, 0.5–1.3(–3.2) mm.

Seeds

(1–)1.2–2(–2.3) × 0.7–1(–1.3) mm.

2n

= 26.

Thelypodium integrifolium subsp. affine

Thelypodium integrifolium

Phenology Flowering Jun–Oct.
Habitat Desert flats, shrub communities, alkaline grounds
Elevation 600-1800 m (2000-5900 ft)
Distribution
from FNA
CA; NV; UT
[BONAP county map]
from FNA
AZ; CA; CO; ID; MT; ND; NE; NM; NV; OR; SD; UT; WA; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 5 (5 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals white; fruiting pedicels whitish, stout, (5-) 6-11(-13) mm
→ 2
1. Petals usually lavender to purple, rarely white; fruiting pedicels not whitish, slender and 2-10 (-13) mm, or stout and 2-5(-6) mm
→ 3
2. Petals (5-)6-9(-10) mm; fruiting pedicels horizontal to divaricate, (5-)6-9(-13) mm; fruits (1.4-)1.9-3.7(-4.5) cm; gynophores (0.5-)1-3 mm; se California, s Nevada, sw Utah.
subsp. affine
2. Petals (7.5-)8-11(-13) mm; fruiting pedicels usually horizontal to reflexed, rarely divaricate, (6-)7-11(-13) mm; fruits (2.2-) 3.5-6.5(-8) cm; gynophores (1-)1.5-4 (-5.5) mm; Grand Canyon, Colorado River, Arizona.
subsp. longicarpum
3. Fruiting pedicels often slender, sometimes stout, not or slightly flattened at base, (4-)6- 10 (-13) mm.
subsp. integrifolium
3. Fruiting pedicels stout, strongly flattened at base, 2-5(-6) mm
→ 4
4. Racemes often strongly congested, central rachises (0.6-)1.2-7(-9) cm; gynophores stout, 0.5-1(-3) mm; e California, Nevada, s Oregon, nw Utah.
subsp. complanatum
4. Racemes somewhat congested, central rachises (3-)8-20(-28) cm; gynophores slender, (0.5-)0.8-3(-4) mm; ne Arizona, w Colorado, nw New Mexico, e Utah.
subsp. gracilipes
Source FNA vol. 7, p. 733. FNA vol. 7, p. 732.
Parent taxa Brassicaceae > tribe Thelypodieae > Thelypodium > Thelypodium integrifolium Brassicaceae > tribe Thelypodieae > Thelypodium
Sibling taxa
T. integrifolium subsp. complanatum, T. integrifolium subsp. gracilipes, T. integrifolium subsp. integrifolium, T. integrifolium subsp. longicarpum
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
Subordinate taxa
T. integrifolium subsp. affine, T. integrifolium subsp. complanatum, T. integrifolium subsp. gracilipes, T. integrifolium subsp. integrifolium, T. integrifolium subsp. longicarpum
Synonyms T. affine, Pleurophragma rhomboideum, T. integrifolium var. affine, T. rhomboideum Pachypodium integrifolium, Pleurophragma integrifolium
Name authority (Greene) Al-Shehbaz: Contr. Gray Herb. 204: 110. (1973) (Nuttall) Endlicher: in W. G. Walpers, Repert. Bot. Syst. 1: 172. (1842)
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