Thelypodium eucosmum |
Thelypodium |
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arrow-leaf thelypody, world thelypody |
thelypody |
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Habit | Biennials or perennials; (short-lived); glaucous, glabrous (except petioles). | Biennials, perennials, or, rarely, annuals; not scapose; glabrous or pubescent. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | branched distally, 2–10 dm. |
(simple or few to several from base), usually erect, rarely decumbent, branched basally and/or distally, (glabrous or pubescent). |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate or not, petiolate, blade margins usually entire, dentate, lyrate or pinnately lobed, rarely laciniate; cauline petiolate or sessile, blade (base cuneate, attenuate, auriculate, sagittate, or amplexicaul), margins often entire, sometimes dentate or pinnately lobed. |
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Basal leaves | petiole (0.9–)1.4–3(–4.5) cm, ciliate; blade usually oblanceolate to oblong or lanceolate, rarely ovate or elliptic, (2.8–)3.5–8.8(–11) cm × (7–)10–25(–35) mm, margins often entire or repand, sometimes sinuate. |
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Cauline leaves | (ascending); sessile; blade lanceolate to oblong, 1.8–4.5(–6) cm × 5–16(–24) mm, (base amplexicaul to strongly auriculate), margins entire. |
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Racemes | lax, slightly elongated in fruit, (flower buds oblong-linear). |
(corymbose, dense or lax), usually slightly to considerably elongated in fruit (sometimes not elongated in T. integrifolium). |
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Flowers | sepals erect, linear-oblong, 5–7(–8) × (0.8–)1–1.5(–1.8) mm; petals dark purple, spatulate to oblanceolate, (6.6–)7.5–10(–11.5) × 1–1.8(–2) mm, margins not crisped, claw differentiated from blade, [slender, (3–)3.5–5(–5.5) mm, narrowest at base]; nectar glands confluent, subtending bases of stamens; filaments subequal, (4.5–)6–9(–10) mm; anthers exserted, linear to narrowly oblong, 2.5–4(–4.5) mm, circinately coiled; gynophore (1–)2.5–6(–7.5) mm. |
sepals usually erect or ascending, rarely spreading to reflexed, ovate to oblong, linear, lanceolate, or oblanceolate, lateral pair slightly saccate or not basally; petals (erect or spreading), white, lavender, or purple, spatulate to obovate, or oblanceolate to linear, (margins crisped or not), claw differentiated or not from blade, (apex rounded); stamens subequal or tetradynamous, (exserted or included); filaments (erect or spreading, usually distinct, very rarely median ones united), not dilated basally; anthers usually linear to linear-oblong, rarely oblong or ovate, (sometimes apiculate, often circinately coiled after dehiscence); nectar glands confluent and subtending bases of stamens, or 2 or 4 and lateral. |
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Fruiting pedicels | horizontal to divaricate, usually straight, rarely slightly incurved, slender, (2.5–)3–5.5(–6.5) mm, slightly flattened at base. |
usually horizontal, erect to ascending, or divaricate, rarely reflexed, slender or stout, (flattened or not basally, glabrous). |
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Fruits | divaricate to ascending, torulose, straight or slightly incurved, terete, (2–)2.4–5(–6.5) cm × 0.7–1(–1.3) mm; ovules 44–58 per ovary; style usually cylindrical, rarely subclavate, 0.5–1.5(–2) mm. |
stipitate, linear, torulose or smooth, terete, slightly 4-angled, or flattened; valves each with prominent midvein, glabrous; replum rounded; septum complete; ovules 12–128 per ovary; style distinct, (often cylindrical, rarely subclavate or subconical); stigma capitate, entire. |
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Seeds | 0.7–1.5 × 0.5–0.8 mm. |
uniseriate, plump or flattened, not winged, usually oblong, rarely ovate; seed coat (minutely reticulate), not mucilaginous when wetted; cotyledons oblique, rarely incumbent or accumbent. |
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x | = 13. |
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Thelypodium eucosmum |
Thelypodium |
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Phenology | Flowering May–Jul. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Shady slopes and canyons, pinyon-juniper and oak woodland communities, stream beds, streamsides | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 700-1000 m (2300-3300 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
OR
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w North America; n Mexico; wc North America |
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Discussion | Of conservation concern. Thelypodium eucosmum is known only from Baker, Grant, and Wheeler counties. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 16 (16 in the flora). As recognized herein and by recent authors (e.g., I. A. Al-Shehbaz 1973; R. C. Rollins 1993), Thelypodium is somewhat heterogeneous and the segregate Stanleyella might merit recognition, as by E. B. Payson (1923). Thelypodium has erect sepals, petals, and stamens, terete fruits, prominently veined fruit septa, and, often, cylindrical styles. In contrast, species of Stanleylla have spreading sepals, petals, and stamens, flattened fruits, veinless fruit septa, and clavate, subclavate, or subconical styles. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 731. | FNA vol. 7, p. 728. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Thelypodieae > Thelypodium | Brassicaceae > tribe Thelypodieae | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Pachypodium, Pleurophragma, Stanleyella | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1(1,1): 175. (1895) | Endlicher: Gen. Pl. 11: 876. (1839) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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