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short-pod thelypodium, short-pod thelypody

Wright's stanleyella, Wright's thelypody

Habit Biennials; often glaucous, glabrous or pubescent. Biennials; slightly glaucous, glabrous.
Stems

branched, (1.3–)3.4–8.3(–12) dm, (somewhat glaucous, glabrous throughout or pubescent proximally).

branched distally, (6.5–)9.5–22.5(–28) dm.

Basal leaves

petiole (1.5–)2–5.2(–9) cm, glabrous or pubescent;

blade usually oblanceolate to spatulate, rarely obovate or lanceolate, 3.3–14(–20) cm × (05–)10–32(–47) mm, margins often pinnately lobed to lyrate, sometimes dentate or entire.

(and proximal cauline): petiole (2–)4.5–8.5(–13.7) cm;

blade often lanceolate, sometimes oblanceolate in outline (lateral lobes oblong to linear or deltate), (6.5–)9.5–22.5(–28) cm × 28–55(–75) mm, margins pinnately lobed or lyrate (lobes entire or denticulate).

Cauline leaves

sessile;

blade linear to lanceolate, smaller distally, (base sagittate, appressed to stem), margins usually entire, rarely dentate.

petiolate;

blade lanceolate to linear-lanceolate, 3.5–8.5(–13.5) cm × 7–14(–23) mm, margins usually entire or dentate, rarely lobed.

Racemes

dense, slightly elongated in fruit, (flower buds oblong).

somewhat lax, considerably elongated in fruit.

Flowers

sepals erect, lanceolate to linear-lanceolate or ovate, (3–)3.5–5(–5.5) × 1–1.5(–2) mm;

petals white, linear to narrowly oblanceolate, 8–12.5(–16) × 0.3–0.5(–1) mm, margins strongly crisped, claw differentiated from blade, (slender, 2.5–4 mm, narrowest at base);

nectar glands surrounding bases of lateral stamens, median glands absent;

filaments subequal or slightly tetradynamous, 2.5–6.5(–10) mm;

anthers exserted, linear, 1.5.–2(–2.5) mm, slightly circinately coiled, (apiculate);

gynophore (0.5–)1–2(–5) mm.

sepals spreading or ascending oblong to linear, (3–)4–6(–7) × (0.8–)1–1.3(–1.5) mm;

petals usually white, rarely lavender, linear or oblong, 4–7.5(–9) × 1–1.8(–2) mm, margins not crisped, claw not differentiated from blade (base sometimes clawlike, to 2 mm, widest at base);

nectar glands confluent;

filaments equal, (3–)3.5–6.5(–8.5) mm;

anthers (1–)1.5–2.5(–3) mm, circinately coiled;

gynophore 0.2–2(–5) mm.

Fruiting pedicels

horizontal to divaricate, straight or slightly curved, stout, 1–2(–2.5) mm, flattened at base.

horizontal to divaricate, sometimes reflexed, straight or curved, sometimes secund, slender, (5–)7–13(–17) mm, slightly flattened at base.

Fruits

divaricate to ascending, torulose, often straight, sometimes slightly curved, terete, (0.8–)1.2–2.7(–3) cm × 1–1.5(–2) mm;

ovules 12–26 per ovary;

style cylindrical, (0.2–)0.5–1(–2.5) mm.

horizontal or reflexed, torulose, straight to somewhat curved, flattened, (2.5–)3.8–7.4(–9) cm × 1–1.2(–1.5) mm, (replum not constricted between seeds);

ovules 76–128 per ovary;

style usually clavate to subclavate, rarely cylindrical, (0.5–)0.8–2(–3) mm.

Seeds

(plump), (1.3–)1.5–2 × 0.8–1(–1.3) mm.

0.7–1.3(–1.5) × 0.5–0.8 mm.

Thelypodium brachycarpum

Thelypodium wrightii

Phenology Flowering Apr–Aug. Flowering (Mar-)Jun–Oct.
Habitat Strongly alkaline meadows and desert flats Rock crevices, shady slopes, rocky hillsides, canyons, stream banks, creek beds, pinyon-juniper communities, oak woodlands
Elevation 600-2300 m (2000-7500 ft) 1200-2300 m (3900-7500 ft)
Distribution
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Hidalgo)
[WildflowerSearch map]
[BONAP county map]
Discussion

Thelypodium brachycarpum is known in California from Napa, Shasta, and Siskiyou counties and in Oregon from Klamath County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Study of extensive material that has accumulated during the past three decades reveals that the differences between subsp. oklahomensis and subsp. wrightii, which are based solely on variations in gynophore and fruit lengths, do not hold as clearly as previously thought (I. A. Al-Shehbaz 1973; R. C. Rollins 1993).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 730. FNA vol. 7, p. 738.
Parent taxa Brassicaceae > tribe Thelypodieae > Thelypodium Brassicaceae > tribe Thelypodieae > Thelypodium
Sibling taxa
T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum, T. wrightii
T. brachycarpum, T. crispum, T. eucosmum, T. flexuosum, T. howellii, T. integrifolium, T. laciniatum, T. laxiflorum, T. milleflorum, T. paniculatum, T. repandum, T. rollinsii, T. sagittatum, T. stenopetalum, T. texanum
Synonyms Thelypodiopsis brachycarpa, Thelypodiopsis brachypoda Stanleyella wrightii, T. wrightii subsp. oklahomensis
Name authority Torrey: in C. Wilkes et al., U.S. Expl. Exped. 17(2): 231, plate 1. (1874) A. Gray: Smithsonian Contr. Knowl. 3(5): 7. (1852)
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