Thelypodium brachycarpum |
Thelypodium |
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short-pod thelypodium, short-pod thelypody |
thelypody |
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Habit | Biennials; often glaucous, glabrous or pubescent. | Biennials, perennials, or, rarely, annuals; not scapose; glabrous or pubescent. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | branched, (1.3–)3.4–8.3(–12) dm, (somewhat glaucous, glabrous throughout or pubescent proximally). |
(simple or few to several from base), usually erect, rarely decumbent, branched basally and/or distally, (glabrous or pubescent). |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate or not, petiolate, blade margins usually entire, dentate, lyrate or pinnately lobed, rarely laciniate; cauline petiolate or sessile, blade (base cuneate, attenuate, auriculate, sagittate, or amplexicaul), margins often entire, sometimes dentate or pinnately lobed. |
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Basal leaves | petiole (1.5–)2–5.2(–9) cm, glabrous or pubescent; blade usually oblanceolate to spatulate, rarely obovate or lanceolate, 3.3–14(–20) cm × (05–)10–32(–47) mm, margins often pinnately lobed to lyrate, sometimes dentate or entire. |
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Cauline leaves | sessile; blade linear to lanceolate, smaller distally, (base sagittate, appressed to stem), margins usually entire, rarely dentate. |
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Racemes | dense, slightly elongated in fruit, (flower buds oblong). |
(corymbose, dense or lax), usually slightly to considerably elongated in fruit (sometimes not elongated in T. integrifolium). |
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Flowers | sepals erect, lanceolate to linear-lanceolate or ovate, (3–)3.5–5(–5.5) × 1–1.5(–2) mm; petals white, linear to narrowly oblanceolate, 8–12.5(–16) × 0.3–0.5(–1) mm, margins strongly crisped, claw differentiated from blade, (slender, 2.5–4 mm, narrowest at base); nectar glands surrounding bases of lateral stamens, median glands absent; filaments subequal or slightly tetradynamous, 2.5–6.5(–10) mm; anthers exserted, linear, 1.5.–2(–2.5) mm, slightly circinately coiled, (apiculate); gynophore (0.5–)1–2(–5) mm. |
sepals usually erect or ascending, rarely spreading to reflexed, ovate to oblong, linear, lanceolate, or oblanceolate, lateral pair slightly saccate or not basally; petals (erect or spreading), white, lavender, or purple, spatulate to obovate, or oblanceolate to linear, (margins crisped or not), claw differentiated or not from blade, (apex rounded); stamens subequal or tetradynamous, (exserted or included); filaments (erect or spreading, usually distinct, very rarely median ones united), not dilated basally; anthers usually linear to linear-oblong, rarely oblong or ovate, (sometimes apiculate, often circinately coiled after dehiscence); nectar glands confluent and subtending bases of stamens, or 2 or 4 and lateral. |
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Fruiting pedicels | horizontal to divaricate, straight or slightly curved, stout, 1–2(–2.5) mm, flattened at base. |
usually horizontal, erect to ascending, or divaricate, rarely reflexed, slender or stout, (flattened or not basally, glabrous). |
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Fruits | divaricate to ascending, torulose, often straight, sometimes slightly curved, terete, (0.8–)1.2–2.7(–3) cm × 1–1.5(–2) mm; ovules 12–26 per ovary; style cylindrical, (0.2–)0.5–1(–2.5) mm. |
stipitate, linear, torulose or smooth, terete, slightly 4-angled, or flattened; valves each with prominent midvein, glabrous; replum rounded; septum complete; ovules 12–128 per ovary; style distinct, (often cylindrical, rarely subclavate or subconical); stigma capitate, entire. |
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Seeds | (plump), (1.3–)1.5–2 × 0.8–1(–1.3) mm. |
uniseriate, plump or flattened, not winged, usually oblong, rarely ovate; seed coat (minutely reticulate), not mucilaginous when wetted; cotyledons oblique, rarely incumbent or accumbent. |
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x | = 13. |
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Thelypodium brachycarpum |
Thelypodium |
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Phenology | Flowering Apr–Aug. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Strongly alkaline meadows and desert flats | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 600-2300 m (2000-7500 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA; OR
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w North America; n Mexico; wc North America |
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Discussion | Thelypodium brachycarpum is known in California from Napa, Shasta, and Siskiyou counties and in Oregon from Klamath County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 16 (16 in the flora). As recognized herein and by recent authors (e.g., I. A. Al-Shehbaz 1973; R. C. Rollins 1993), Thelypodium is somewhat heterogeneous and the segregate Stanleyella might merit recognition, as by E. B. Payson (1923). Thelypodium has erect sepals, petals, and stamens, terete fruits, prominently veined fruit septa, and, often, cylindrical styles. In contrast, species of Stanleylla have spreading sepals, petals, and stamens, flattened fruits, veinless fruit septa, and clavate, subclavate, or subconical styles. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 730. | FNA vol. 7, p. 728. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Thelypodieae > Thelypodium | Brassicaceae > tribe Thelypodieae | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Thelypodiopsis brachycarpa, Thelypodiopsis brachypoda | Pachypodium, Pleurophragma, Stanleyella | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Torrey: in C. Wilkes et al., U.S. Expl. Exped. 17(2): 231, plate 1. (1874) | Endlicher: Gen. Pl. 11: 876. (1839) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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