Thelypodium |
Thelypodium stenopetalum |
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thelypody |
slender-petal thelypodium, slenderpetal thelypody |
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Habit | Biennials, perennials, or, rarely, annuals; not scapose; glabrous or pubescent. | Biennials; glaucous, glabrous (except petioles). | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | (simple or few to several from base), usually erect, rarely decumbent, branched basally and/or distally, (glabrous or pubescent). |
(often decumbent), branched basally, sometimes also distally, (2.6–)3–8(–9) dm. |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate or not, petiolate, blade margins usually entire, dentate, lyrate or pinnately lobed, rarely laciniate; cauline petiolate or sessile, blade (base cuneate, attenuate, auriculate, sagittate, or amplexicaul), margins often entire, sometimes dentate or pinnately lobed. |
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Basal leaves | petiole 1–4.5(–6) cm, ciliate; blade often oblong or lanceolate, sometimes ovate to spatulate, 3.8–15(–18) cm × 15–35(–42) mm, margins usually entire, rarely repand. |
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Cauline leaves | (ascending); sessile; blade usually lanceolate to oblong, rarely linear to linear-lanceolate, (1.3–)1.6–4.8(–6) cm × (3–)5–9(–15) mm, (base usually amplexicaul to sagittate, rarely auriculate), margins entire. |
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Racemes | (corymbose, dense or lax), usually slightly to considerably elongated in fruit (sometimes not elongated in T. integrifolium). |
lax, elongated in fruit, (flower buds oblong-linear). |
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Flowers | sepals usually erect or ascending, rarely spreading to reflexed, ovate to oblong, linear, lanceolate, or oblanceolate, lateral pair slightly saccate or not basally; petals (erect or spreading), white, lavender, or purple, spatulate to obovate, or oblanceolate to linear, (margins crisped or not), claw differentiated or not from blade, (apex rounded); stamens subequal or tetradynamous, (exserted or included); filaments (erect or spreading, usually distinct, very rarely median ones united), not dilated basally; anthers usually linear to linear-oblong, rarely oblong or ovate, (sometimes apiculate, often circinately coiled after dehiscence); nectar glands confluent and subtending bases of stamens, or 2 or 4 and lateral. |
sepals erect, linear to linear-oblong, (6–)6.5–9(–10) × 1–1.5(–1.8) mm; petals usually lavender, rarely white, linear, (8–)9.5–15(–16.5) × 0.3–0.5(–0.8) mm, margins crisped between blade and claw, claw differentiated from blade, [slender, 4–6.5(–8.5) mm, narrowest at base]; nectar glands confluent, subtending bases of stamens; filaments slightly tetradynamous, median pairs (7–)8–12.5(–14) mm, lateral pair (5.5–)7–11(–12) mm; anthers exserted, linear to narrowly oblong, 3.5–5(–6) mm, circinately coiled; gynophore stout, 0.5–3.5(–5) mm. |
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Fruiting pedicels | usually horizontal, erect to ascending, or divaricate, rarely reflexed, slender or stout, (flattened or not basally, glabrous). |
usually ascending to divaricate-ascending, rarely horizontal or divaricate, usually straight, rarely slightly incurved, stout, (3.5–)4–8 mm, not flattened at base. |
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Fruits | stipitate, linear, torulose or smooth, terete, slightly 4-angled, or flattened; valves each with prominent midvein, glabrous; replum rounded; septum complete; ovules 12–128 per ovary; style distinct, (often cylindrical, rarely subclavate or subconical); stigma capitate, entire. |
usually divaricate-ascending to ascending, rarely divaricate, slightly torulose, straight or slightly incurved, terete or slightly 4-angled, (2.2–)2.8–5(–6.3) cm × 1–1.5(–1.8) mm; ovules 50–82 per ovary; style cylindrical, 1–2(–2.5) mm. |
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Seeds | uniseriate, plump or flattened, not winged, usually oblong, rarely ovate; seed coat (minutely reticulate), not mucilaginous when wetted; cotyledons oblique, rarely incumbent or accumbent. |
1–1.3(–1.5) × 0.7–0.8 mm. |
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x | = 13. |
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Thelypodium |
Thelypodium stenopetalum |
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Phenology | Flowering May–Aug. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Alkaline meadows and flats | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 1900-2100 m (6200-6900 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
w North America; n Mexico; wc North America |
CA |
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Discussion | Species 16 (16 in the flora). As recognized herein and by recent authors (e.g., I. A. Al-Shehbaz 1973; R. C. Rollins 1993), Thelypodium is somewhat heterogeneous and the segregate Stanleyella might merit recognition, as by E. B. Payson (1923). Thelypodium has erect sepals, petals, and stamens, terete fruits, prominently veined fruit septa, and, often, cylindrical styles. In contrast, species of Stanleylla have spreading sepals, petals, and stamens, flattened fruits, veinless fruit septa, and clavate, subclavate, or subconical styles. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Thelypodium stenopetalum is known from Bear Valley, San Bernardino County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 728. | FNA vol. 7, p. 737. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Pachypodium, Pleurophragma, Stanleyella | Thelypodiopsis stenopetala | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Endlicher: Gen. Pl. 11: 876. (1839) | S. Watson: Proc. Amer. Acad. Arts 22: 468. (1887) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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