Thelypodium |
Thelypodium flexuosum |
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thelypody |
nodding thelypodium, nodding thelypody, spreading thelypodium, spreading thelypody, zigzag thelypody |
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Habit | Biennials, perennials, or, rarely, annuals; not scapose; glabrous or pubescent. | Perennials; (caudex woody, covered with persistent petiolar remains); somewhat glaucous, glabrous throughout. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | (simple or few to several from base), usually erect, rarely decumbent, branched basally and/or distally, (glabrous or pubescent). |
(often subdecumbent), branched basally, (flexuous), 1.5–5.6(–8.5) dm. |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate or not, petiolate, blade margins usually entire, dentate, lyrate or pinnately lobed, rarely laciniate; cauline petiolate or sessile, blade (base cuneate, attenuate, auriculate, sagittate, or amplexicaul), margins often entire, sometimes dentate or pinnately lobed. |
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Basal leaves | petiole 1–6.5(–13) cm; blade often lanceolate, sometimes oblong or oblanceolate, (2–)3.5–16.5(–20.5) cm × (5–)10–25(–45) mm, margins entire. |
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Cauline leaves | (ascending); sessile; blade lanceolate to linear, 1–7(–11) cm × 2–7(–14) mm, (base sagittate to somewhat amplexicaul), margins entire. |
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Racemes | (corymbose, dense or lax), usually slightly to considerably elongated in fruit (sometimes not elongated in T. integrifolium). |
(few-flowered, corymbose), elongated in fruit, (flower buds oblong). |
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Flowers | sepals usually erect or ascending, rarely spreading to reflexed, ovate to oblong, linear, lanceolate, or oblanceolate, lateral pair slightly saccate or not basally; petals (erect or spreading), white, lavender, or purple, spatulate to obovate, or oblanceolate to linear, (margins crisped or not), claw differentiated or not from blade, (apex rounded); stamens subequal or tetradynamous, (exserted or included); filaments (erect or spreading, usually distinct, very rarely median ones united), not dilated basally; anthers usually linear to linear-oblong, rarely oblong or ovate, (sometimes apiculate, often circinately coiled after dehiscence); nectar glands confluent and subtending bases of stamens, or 2 or 4 and lateral. |
sepals erect, oblong, 3–4(–4.5) × 1–1.5(–1.7) mm; petals lavender or white, often spatulate, sometimes oblanceolate or obovate, 6–9(–10) × (1.5–)2–3(–3.5) mm, margins not crisped, claw strongly differentiated from blade, (slender, 2–3.5(–4) mm, narrowest at base); nectar glands lateral, median glands absent; filaments tetradynamous, median pairs 3–4(–5) mm, lateral pair 2–3.5(–4) mm; anthers included, oblong, 1–2(–2.5) mm, not circinately coiled; gynophore (stout), 0.5–1 mm. |
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Fruiting pedicels | usually horizontal, erect to ascending, or divaricate, rarely reflexed, slender or stout, (flattened or not basally, glabrous). |
usually horizontal to divaricate, rarely divaricate-ascending, straight or slightly curved upward, slender, (2.5–)4–9(–16) mm, slightly flattened at base. |
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Fruits | stipitate, linear, torulose or smooth, terete, slightly 4-angled, or flattened; valves each with prominent midvein, glabrous; replum rounded; septum complete; ovules 12–128 per ovary; style distinct, (often cylindrical, rarely subclavate or subconical); stigma capitate, entire. |
erect to ascending, torulose, slightly incurved or straight, terete, 1–2.5(–4.2) cm × 0.8–1(–1.5) mm; ovules 12–30 per ovary; style cylindrical, (0.3–)1–2(–3) mm. |
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Seeds | uniseriate, plump or flattened, not winged, usually oblong, rarely ovate; seed coat (minutely reticulate), not mucilaginous when wetted; cotyledons oblique, rarely incumbent or accumbent. |
(1–)1.3–1.5 × 0.5–8(–1) mm. |
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x | = 13. |
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2n | = 26. |
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Thelypodium |
Thelypodium flexuosum |
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Phenology | Flowering Apr–Jun. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Often tangled among woody shrubs in moderately to strongly alkaline sandy loam or clay, open deserts | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 1000-2400 m (3300-7900 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
w North America; n Mexico; wc North America |
CA; ID; NV; OR
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Discussion | Species 16 (16 in the flora). As recognized herein and by recent authors (e.g., I. A. Al-Shehbaz 1973; R. C. Rollins 1993), Thelypodium is somewhat heterogeneous and the segregate Stanleyella might merit recognition, as by E. B. Payson (1923). Thelypodium has erect sepals, petals, and stamens, terete fruits, prominently veined fruit septa, and, often, cylindrical styles. In contrast, species of Stanleylla have spreading sepals, petals, and stamens, flattened fruits, veinless fruit septa, and clavate, subclavate, or subconical styles. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 728. | FNA vol. 7, p. 731. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Pachypodium, Pleurophragma, Stanleyella | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Endlicher: Gen. Pl. 11: 876. (1839) | B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1(1,1): 175. (1895) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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