Thelypodium |
Thelypodium crispum |
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thelypody |
crisp thelypody, crisped thelypody, wavy leaf thelypodium |
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Habit | Biennials, perennials, or, rarely, annuals; not scapose; glabrous or pubescent. | Biennials or perennials; (short-lived); often glaucous, glabrous or pubescent. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | (simple or few to several from base), usually erect, rarely decumbent, branched basally and/or distally, (glabrous or pubescent). |
branched, (0.9–)1.3–7(–12) dm, (somewhat glaucous, glabrous throughout or pubescent basally). |
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Leaves | basal and cauline; petiolate or sessile; basal rosulate or not, petiolate, blade margins usually entire, dentate, lyrate or pinnately lobed, rarely laciniate; cauline petiolate or sessile, blade (base cuneate, attenuate, auriculate, sagittate, or amplexicaul), margins often entire, sometimes dentate or pinnately lobed. |
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Basal leaves | petiole (0.7–)2–8.5 cm, ciliate; blade usually oblanceolate to spatulate, rarely obovate or lanceolate, (1.5–)2.2–15(–25) cm × (6–)10–35(–50) mm, margins often pinnately lobed to lyrate, sometimes dentate or entire. |
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Cauline leaves | sessile; blade linear to lanceolate, smaller distally, (base sagittate or auriculate, appressed to stem), margins usually entire, rarely dentate. |
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Racemes | (corymbose, dense or lax), usually slightly to considerably elongated in fruit (sometimes not elongated in T. integrifolium). |
dense, elongated in fruit, (flower buds narrowly oblong). |
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Flowers | sepals usually erect or ascending, rarely spreading to reflexed, ovate to oblong, linear, lanceolate, or oblanceolate, lateral pair slightly saccate or not basally; petals (erect or spreading), white, lavender, or purple, spatulate to obovate, or oblanceolate to linear, (margins crisped or not), claw differentiated or not from blade, (apex rounded); stamens subequal or tetradynamous, (exserted or included); filaments (erect or spreading, usually distinct, very rarely median ones united), not dilated basally; anthers usually linear to linear-oblong, rarely oblong or ovate, (sometimes apiculate, often circinately coiled after dehiscence); nectar glands confluent and subtending bases of stamens, or 2 or 4 and lateral. |
sepals erect to ascending, oblong to linear-oblong, (3–)3.5–6(–8) × (0.8–)1–1.8(–2) mm; petals white to lavender, linear to narrowly oblanceolate, (6–)6.5–11(–14.5) × 0.5–0.7(–1) mm, margins strongly crisped, claw differentiated from blade, (slender, 2–4 mm, narrowest at base); nectar glands surrounding bases of lateral stamens, median glands absent; filaments subequal, (2.5–)4.5–8.5(–10.5) mm; anthers exserted, linear to narrowly oblong, 2–3.5(–4.5) mm, circinately coiled, (apiculate); gynophore 0.5–1.5(–3.5) mm. |
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Fruiting pedicels | usually horizontal, erect to ascending, or divaricate, rarely reflexed, slender or stout, (flattened or not basally, glabrous). |
erect to erect-ascending, (appressed to rachis at least basally), straight or slightly curved, slender, (1.5–)2–5(–10) mm, not flattened at base. |
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Fruits | stipitate, linear, torulose or smooth, terete, slightly 4-angled, or flattened; valves each with prominent midvein, glabrous; replum rounded; septum complete; ovules 12–128 per ovary; style distinct, (often cylindrical, rarely subclavate or subconical); stigma capitate, entire. |
divaricate to ascending, torulose, straight or slightly curved, terete, (0.6–)1–2.5(–4.2) cm × 0.7–1(–1.8) mm; ovules 22–50 per ovary; style cylindrical, 0.5–1.5(–2.5) mm. |
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Seeds | uniseriate, plump or flattened, not winged, usually oblong, rarely ovate; seed coat (minutely reticulate), not mucilaginous when wetted; cotyledons oblique, rarely incumbent or accumbent. |
(flattened), 1–1.5(–1.7) × 0.5–0.7(–1) mm. |
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x | = 13. |
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2n | = 26. |
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Thelypodium |
Thelypodium crispum |
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Phenology | Flowering Jun–Aug. | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Alkaline meadows and desert flats, mineralized ground near hot springs, desert shrub communities | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 1200-3000(-3200) m (3900-9800(-10500) ft) | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
w North America; n Mexico; wc North America |
CA; NV
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Discussion | Species 16 (16 in the flora). As recognized herein and by recent authors (e.g., I. A. Al-Shehbaz 1973; R. C. Rollins 1993), Thelypodium is somewhat heterogeneous and the segregate Stanleyella might merit recognition, as by E. B. Payson (1923). Thelypodium has erect sepals, petals, and stamens, terete fruits, prominently veined fruit septa, and, often, cylindrical styles. In contrast, species of Stanleylla have spreading sepals, petals, and stamens, flattened fruits, veinless fruit septa, and clavate, subclavate, or subconical styles. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 728. | FNA vol. 7, p. 730. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | Pachypodium, Pleurophragma, Stanleyella | Thelypodiopsis crispa, T. brachycarpum var. crispum | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Endlicher: Gen. Pl. 11: 876. (1839) | Greene ex Payson: Ann. Missouri Bot. Gard. 9: 264. (1923) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |